(C) _Visceral Arches._--The skeleton of the visceral arches consists
essentially of a series of half-hoops of cartilage, each divided in the
adult into a number of segments and connected with its fellow by a median
ventral cartilage. The skeleton of arches I. and II. (mandibular and
hyoidean) undergoes modifications of special interest (figs. 14 and 15).
The lower portion of the mandibular arch becomes greatly thickened to
support the lower or hinder edge of the mouth. It forms the primitive
lower jaw or "Meckel's cartilage." Dorsal to this an outgrowth arises from
the anterior face of the arch which supports the upper or anterior margin
of the mouth: it is the primitive upper jaw or palato-pterygoquadrate
cartilage. The portion of the arch dorsal to the palato-pterygo-quadrate
outgrowth may form the suspensorial apparatus of the lower jaw, being
fused with the cranium at its upper end. This relatively primitive
con-arrangement (_protostylic_, as it may be termed) occurs in Dipneusti
among fishes (cf. fig. 14). More usually this dorsal part of the
mandibular arch becomes reduced, its place being occupied by a ligament
(pre-spiracular) uniting the jaw apparatus to the chondrocranium, the
upper jaw being also attached to the chondrocranium by the ethmopalatine
ligament situated more anteriorly. The main attachment, however, of the
jaws to the chondrocranium in such a case, as holds for the majority of
fishes, is through the enlarged dorsal segment of the hyoid arch
(hyomandibular) which articulates at its dorsal end with the
chondrocranium, while its ventral end is attached to the hinge region of
the jaw by stout ligamentous bands. A skull in which the jaws are
suspended in this manner is termed a hyostylic skull (e.g. _Scyllium_ in
fig. 15).
[Illustration: FIG. 16.--Fore-limb of _Ceratodus_.]
In _Notidanus_ (fig. 15, B) there is a large direct articulation of
the upper jaw to the chondrocranium in addition to the indirect one
through the hyomandibular: such a skull is amphistylic. In
_Heterodontus_ the upper jaw is firmly bound to the cranium throughout
its length, while in Holocephali (fig. 15, C) complete fusion has
taken place, so that the lower jaw appears to articulate directly with
the cranium ("auto stylic" condition). In Dipneusti[29] (_Lepidosiren_
and _Protopterus_) the cartilaginous upper jaw never develops (except
in its hinder quadrate portion) beyond the condition of a faint
rudiment, owing doubtless to its being replaced functionally by
precociously developed bone.
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