subopercular and opercular.
[Illustration: From Wiedersheim, _Verg. Anat. der Wirbeltiere_, by
permission of Gustav Fischer.
FIG. 25.--Complete Skull of Salmon from left side.
art, Articular. op, Opercular.
branchiost, Branchiostegal. pal, Palatine.
dent, Dentary. par, Parietal.
epiot, Epiotic. pmx, Premaxilla.
eth, Dermal ethmoid. preop, Preopercular.
fr, Frontal. pt, Pterygoid.
hyom, Hyomandibular. pter, Pterotic.
intop, Interopercular. Quad, Quadrate.
Jug, Jugal. socc, Supraoccipital.
mpt, Mesopterygoid. sphot, Sphenotic.
mtpt, Metapterygoid. subop, Subopercular.
mx, Maxilla. sympl, Symplectic.
nas, Nasal. Zunge, Tongue.]
In addition to the bones already enumerated there is present a ring of
circumorbital bones, a preopercular, behind and external to the
hyomandibular and quadrate, and squamosal, external to the hinder end of
the auditory capsule.
In the salmon, pike, and various other Teleosts, extensive regions of
the chondrocranium persist in the adult, while in others (e.g. the
cod) the replacement by bone is practically complete. Bony elements
may be developed in addition to those noticed in the salmon.
In the sturgeon the chondrocranium is ensheathed by numerous membrane
bones, but cartilage bones are absent. In the Crossopterygians[35] the
chondrocranium persists to a great extent in the adult, but portions
of it are replaced by cartilage bones--the most interesting being a
large sphenethmoid like that of the frog. Numerous membrane bones
cover the chondrocranium externally. In the Dipneusti[36] the
chondrocranium is strengthened in the adult by numerous bones. One of
the most characteristic is the great palatopterygoid bone which
develops very early by the spreading of ossification backwards from
the tooth bases, and whose early development probably accounts for the
non-development of the palatopterygoid cartilage.
_Appendicular Skeleton._--The primitive pectoral girdle, which in the
Dipneusti is strengthened by a sheath of bone, becomes in the
Teleostomes reduced in size (small scapula and coracoid bones) and
replaced functionally by a secondary shoulder girdle formed of
superficially placed membrane bones (supraclavicular and cleithrum or
"clavicle," with, in addition in certain cases, an infraclavicular and
one or two postclavicular elements), and connected at its dorsal end
with the skull by a post-temporal bone.
The pelvic girdle is in Teleostomes completely absent as a rule.
The skeleton of the free limb undergoes ossification to a less or
greater extent in the Teleostomes.
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