Hertwig sums up his conclusion in the statement that ontogenetic stages
give us, therefore, a greatly changed picture of the phylogenetic series
of adult ancestors. “The two correspond not according to their actual
contents but only as to their form.” Hertwig also repeats His’s idea,
that the reason that certain kinds of form repeat themselves in the
development of animals with a great constancy depends principally on
this, that they supply the necessary conditions under which alone the
following higher stage of the ontogeny can be formed. The development,
for instance, begins with the division of the egg, because this is the
only way that a one-celled condition can give rise to a many-celled
form. Again, the organs can be formed only when groups of cells have
made a closer union with one another. Thus the gastrula must begin with
the antecedent blastula, etc. Definite forms are, despite all modifying
influences, held to firmly, because by their presence the complicated
end-stages can be reached in the simplest and most suitable way.
Thus Hertwig adopts here a little from one doctrine and there a little
from another, and between his attempt to reinstate the old biogenetic
law of Haeckel, and to adopt a more modern point of view, he brings
together a rather curious collection of statements which are not any too
well coördinated. Take, for example, his description of the relation
between Haeckel’s gastræa and the embryonic gastrula stage. The latter
he maintains is a repetition of the other, but only in form, not in
actual contents. And in another connection we are told that the cause of
this repetition is that the gastrula is the simplest way in which the
later stages can be reached, and, therefore, it has been retained. It
seems to me that Hertwig has undertaken an unnecessary and impossible
task when he attempts to adjust the old recapitulation theory to more
modern standards. His statement that the egg is entirely different from
its amœba prototype is, of course, only the view generally held by all
embryologists. His mystical statement that the embryonic form _repeats
the ancestral adult stage in its form, but not in its contents_, will
scarcely recommend itself as a model of clear thinking. Can we be asked
to believe for instance that a young chick repeats the ancestral adult
fish form but not the contents of the fish?
Public-domain text, read in full here on John Shaqi.
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