In some species there are smaller groups of individuals that are more
like one another than like other individuals of the same species. Such
groups are called varieties, and are often associated with certain
localities, or with a special environment. In the latter case they are
called local varieties. Some of these appear to breed true, not only
when kept under the same conditions, but even when transferred to a new
environment. Others change with the environment. It is not improbable
that the varieties are of a different kind in these two cases, as shown
by their different behavior when put under new and different
surroundings. The variety that owes its peculiarities, not to the
immediate environment, but to some internal condition independent of the
surroundings, is recognized by some biologists as a smaller species.
Such species appear to be commoner in plants than in animals, although
it is possible that this only means that more cases have been found by
the botanists, owing to the greater ease with which plants can be
handled. These smaller species, in contradistinction to the ordinary
Linnæan species, differ from the latter in the smaller amount of
differences between the groups, and probably also in that they freely
interbreed, and leave fertile descendants; but whether this is only on
account of the smaller differences between them than between larger
species, or because of some more fundamental difference in the kind of
variation that gives rise to these two kinds of groups, we do not know.
These smaller species, or constant varieties, as we may call them, may
be looked upon as incipient Linnæan species, which, by further
variations of the same, or of other sorts, may end by giving rise to
true species. A genus composed of several species might be formed in
this way, and then, if each species again broke up into a number of new
groups, each such group would now be recognized as a genus, and the
group of genera would form a family, etc. The process continuing, a
whole class, or order, or even phylum, might be the result of this
process that began in a single species.
But we must look still farther, and inquire whether the start was made
from a single individual, that began to vary, or from a number of
individuals, or even from all the individuals, of a species. If we
suppose the result to depend on some external cause that affects all the
individuals of a species alike, then it might appear that the species,
or at least as many individuals of a species as are affected, will give
the starting-point for the new group. But if the new variation arises
not directly as a response to some change in the surroundings, then it
might appear in one or in a few individuals at a time. Let us consider
what the results might be under these two heads.
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