Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
_Thomomys bottae_ (Eydoux and Gervais, 1836). Mag. de Zool., Paris,
6:23. Type from coast of California, probably near Monterey.
*_Thomomys vetus_ Davis, 1937. Jour. Mamm., 18:156, May 12. Type
from Fossil Lake beds, late Pleistocene, Oregon.
_Thomomys townsendii_ (Bachman, 1839). Jour. Acad. Nat. Sci.
Philadelphia, 8:105. Type probably from near Nampa, Canyon Co.,
Idaho (erroneously given as "Columbia River").
_talpoides_ species-group
*_Thomomys microdon_ Sinclair, 1905. Bull. Dept. Geol. Univ.
California, 4:145-161. Type from Potter Creek Cave, late
Pleistocene, California.
_Thomomys monticola_ J. A. Allen, 1893. Bull. Amer. Mus. Nat. Hist.,
5:48, April 28. Type from Mt. Tallac, 7500 ft., El Dorado Co.,
California.
_Thomomys talpoides_ (Richardson, 1828). Zool. Jour., 3:518. Type
locality fixed at near Fort Carlton (Carlton House),
Saskatchewan River, Saskatchewan, Canada.
_Thomomys mazama_ Merriam, 1897. Proc. Biol. Soc. Washington,
11:214, July 15. Type from Anna Creek, 6000 ft., near Crater
Lake, Mt. Mazama, Klamath Co., Washington.
Tribe GEOMYINI, new tribe
_Genotype._--_Geomys_ Rafinesque, 1817.
_Chronologic and geographic range._--Known from late middle Pliocene
deposits to Recent. The range of living members extends from extreme
southern Manitoba and the southeastern United States south to southern
Panamá, and probably northern Colombia, South America.
_Diagnosis._--Size small to large (condylobasal length of skull 33.0
to 73.0 in adults, including both sexes); sexual dimorphism marked,
sometimes strongly, females being smaller than males, especially in
cranial dimensions; upper incisors invariably grooved, number and
position of grooves varying according to genus; cheek teeth
high-crowned and ever-growing, except in one primitive genus
(_Pliogeomys_); all three lower molars and M1 and M2 monoprismatic,
and elliptical in cross-section in final stages of wear (teeth of
young, subadult, and adult animals); primitive biprismatic patterns
(as known from Recent specimens) occurring only in pre-final stages of
wear (teeth of juveniles only); biprismatic patterns of lower molars
as in _Dikkomys_, and upper molars as in _Pliosaccomys_ (for detailed
description of these patterns, see account beyond of the phylogeny of
the Geomyinae); m3 becoming monoprismatic, anteroposteriorly
compressed and elliptical in cross-section like m1 and m2, but M3
remaining, with rare exceptions (see accounts of _Geomys_ and
_Pappogeomys_ beyond), at least partially biprismatic throughout life,
having one or both lateral inflections usually persisting (with
exceptions) and developing various occlusal shapes (subtriangular,
elongate, obcordate, suborbiculate, or quadriform) but never
elliptical.
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