Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
Of the molariform dentition only the lower premolar and first two
lower molars are known. The enamel investment of p4 is complete, and
would not be subject to interruption at any stage of wear; the two
prisms are joined at their mid-points, and the isthmus of dentine is
relatively broad (as in _Pliosaccomys_) when compared with modern
pocket gophers of this tribe. Also, the re-entrant folds, rather than
having parallel sides, diverge broadly to the sides. The divergence is
especially noticeable in the labial fold. The lower deciduous premolar
would have formed essentially the same enamel pattern with wear as
observed in _Nerterogeomys_ [= _Zygogeomys_] cf. _minor_ (see Hibbard,
1954:fig. 5, A and B) and _Pliosaccomys dubius_ (see Wilson, 1936; pl.
1, fig. 1). Each molar is a single column in the final stages of wear;
pre-final stages are unknown. Anterior and posterior enamel plates are
present on m1 and m2 (m3 has not been recovered). The dentine tracts
of m1 are exposed over a relatively wide surface; therefore, the
enamel plates are distinctly separated. The tracts of dentine of m2
are much narrower than in m1 and the enamel plates are barely
separated at the anterolateral margin of the tooth. Possibly the
enamel band of m2 was continuous in an earlier stage of wear.
The mandible is stout and its general construction not unlike that in
modern geomyines. The capsule at the base of the angular process that
receives the terminal end of the lower incisor is well developed. The
base of the angular processes is preserved, and suggests that the
process was short and decidedly smaller than in living examples of the
tribe. The masseteric ridge is distinct but weakly developed, and not
at all massive as in living pocket gophers. The mental foramen is
immediately anterior, and slightly ventral, to the anterior extension
of the crest. The basitemporal fossa is absent as such, but its
position is marked by a slight depression.
_Specimens examined._--Two rami; nos. 29147 (holotype) and 33446;
several isolated teeth 30194 and 30195, including an upper incisor and
a dp4 (deciduous lower premolar), all from Univ. Michigan Mus. Paleo.
_Referred species._--One.
*_Pliogeomys buisi_ Hibbard, 1954. Papers Michigan Acad. Sci.,
Arts, and Letters, 39:353. Type from Buis local fauna, latest
middle Pliocene, Beaver County, Oklahoma.
Genus =Zygogeomys= Merriam
1895. _Zygogeomys_ Merriam, N. Amer. Fauna, 8:195, January 31.
1942. _Nerterogeomys_ Gazin, Proc. U. S. Nat. Mus., 92:507
(type, _Geomys persimilis_ Hay, 1927).
_Type._--_Zygogeomys trichopus_ Merriam, 1895, from Nahuatzen,
Michoacán.
_Chronologic range._--Late Pliocene (Benson and Curtis Ranch local
faunas, Arizona, and ?Rexroad Formation, Kansas) to Recent.
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