One of the simplest examples is furnished by the development of the
Tubicolar Annelids; but from its very simplicity it appears well
adapted to open the eyes of many who, perhaps, would rather not see,
and it may therefore find a place here. Three years ago I found on the
walls of one of my glasses some small worm-tubes (Fig. 65), the
inhabitants of which bore three pairs of barbate branchial filaments,
and had no operculum. According to this we should have been obliged to
refer them to the genus _Protula._ A few days afterwards one of the
branchial filaments had become thickened at the extremity into a
clavate operculum (Fig. 66), when the animals reminded me, by the
barbate opercular peduncle, of the genus _Filograna,_ only that the
latter possesses two opercula. In three days more, during which a new
pair of branchial filaments had sprouted forth, the opercular peduncle
had lost its lateral filaments (Fig. 67), and the worms had become
_Serpulæ._ Here the supposition at once presents itself that the
primitive tubicolar worm was a _Protula,_—that some of its descendants,
which had already become developed into perfect _Protulæ,_ subsequently
improved themselves by the formation of an operculum which might
protect their tubes from inimical intruders,—and that subsequent
descendants of these latter finally lost the lateral filaments of the
opercular peduncle, which they, like their ancestors, had developed.
What say the schools to this case? Whence and for what purpose, if the
_Serpulæ_ were produced or created as ready-formed species, these
lateral filaments of the opercular peduncle? To allow them to sprout
forth merely for the sake of an invariable plan of structure, even when
they must be immediately retracted again as superfluous, would
certainly be an evidence rather of childish trifling or dictatorial
pedantry, than of infinite wisdom. But no, I am mistaken; from the
beginning of all things the Creator knew, that one day the inquisitive
children of men would grope about after analogies and homologies, and
that Christian naturalists would busy themselves with thinking out his
Creative ideas; at any rate, in order to facilitate the discernment by
the former that the opercular peduncle of the _Serpulæ_ is homologous
with a branchial filament, He allowed it to make a detour in its
development, and pass through the form of a barbate branchial filament.
_The historical record preserved in developmental history is gradually_
EFFACED _as the development strikes into a constantly straighter course
from the egg to the perfect animal, and it is frequently_ SOPHISTICATED
_by the struggle for existence which the free-living larvæ have to
undergo._
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