The changes by which young animals depart from their parents, and the
gradual accumulation of which causes the production of new species,
genera, and families, may occur at an earlier or later period of
life,—in the young state, or at the period of sexual maturity. For the
latter is by no means always, as in the Insecta, a period of repose;
most other animals even then continue to grow and to undergo changes.
(See above, the remarks on the males of the Amphipoda.) Some
variations, indeed, from their very nature, can only occur when the
young animal has attained the adult stage of development. Thus the Sea
Caterpillars (_Polynoë_) at first possess only a few body-segments,
which, during development, gradually increase to a number which is
different in different species, but constant in the same species; now
before a young animal could exceed the number of segments of its
parents, it must of course have attained that number. We may assume a
similar supplementary progress wherever the deviation of the
descendants consists in an addition of new segments and limbs.
_Descendants therefore reach a new goal, either by deviating sooner or
later whilst still on the way towards the form of their parents, or by
passing along this course without deviation, but then, instead of
standing still, advance still farther._
The former mode will have had a predominant action where the posterity
of common ancestors constitutes a group of forms standing upon the same
level in essential features, as the whole of the Amphipoda, Crabs, or
Birds. On the other hand we are led to the assumption of the second
mode of progress, when we seek to deduce from a common original form,
animals some of which agree with young states of others.
In the former case the developmental history of the descendants can
only agree with that of their ancestors up to a certain point at which
their courses separate,—as to their structure in the adult state it
will teach us nothing. _In the second case the entire development of
the progenitors is also passed through by the descendants, and,
therefore, so far as the production of a species depends upon this
second mode of progress, the historical development of the species will
be mirrored in its developmental history._ In the short period of a few
weeks or months, the changing forms of the embryo and larvæ will pass
before us, a more or less complete and more or less true picture of the
transformations through which the species, in the course of untold
thousands of years, has struggled up to its present state.
(Figs. 65 to 67. Young Tubicolar worms, magnified with the simple lens:
65. Without operculum, Protula-stage. 66. With a barbate opercular
peduncle, Filograna-stage; With a naked opercular peduncle,
Serpula-stage. Figs. 65–67. Young Tubicolar worms, magnified with the
simple lens: 65.[1] Without operculum, _Protula_-stage; 66. With a
barbate opercular peduncle, _Filograna_-stage; 67. With a naked
opercular peduncle, _Serpula_-stage.
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