cut leaves a permanent thin mark across the ridges (Fig. 7, _c_),
sometimes without any accompanying puckering; but there is often a
displacement of the ridges on both sides of it, exactly like a "fault" in
stratified rocks. A cut, or other injury that is not a clean incision,
leaves a scar with puckerings on all sides, as in Fig. 7, _a_, making the
ridges at that part undecipherable, even if it does not wholly obliterate
them.
The latest and best investigations on the evolution of the ridges have
been made by Dr. H. Klaatsch.[2] He shows that the earliest appearance in
the Mammalia of structures analogous to ridges is one in which small
eminences occur on the ball of the foot, through which the sweat glands
issue in no particular order. The arrangement of the papillae into rows,
and the accompanying orderly arrangement of the sweat glands, is a
subsequent stage in evolution. The prehensile tail of the Howling Monkey
serves as a fifth hand, and the naked concave part of the tail, with which
it grasps and holds on to boughs, is furnished with ridges arranged
transversely in beautiful order. The numerous drawings of the hands of
monkeys by Allix[3] may be referred to with advantage.
The uses of the ridges are primarily, as I suppose, to raise the mouths of
the ducts, so that the excretions which they pour out may the more easily
be got rid of; and secondarily, in some obscure way, to assist the sense
of touch. They are said to be moulded upon the subcutaneous papillae in
such a manner that the ultimate organs of touch, namely, the Pacinian
bodies, etc.--into the variety of which it is unnecessary here to
enter--are more closely congregated under the bases of the ridges than
under the furrows, and it is easy, on those grounds, to make reasonable
guesses how the ridges may assist the sense of touch. They must
concentrate pressures, that would otherwise be spread over the surface
generally, upon the parts which are most richly supplied with the
terminations of nerves. By their means it would become possible to
neutralise the otherwise dulling effect of a thick protective epidermis.
Their existence in transverse ridges on the inner surface of the
prehensile tails of monkeys admits of easy justification from this point
of view. The ridges so disposed cannot prevent the tail from curling, and
they must add materially to its sensitiveness. They seem to produce the
latter effect on the hands of man, for, as the epidermis thickens under
use within moderate limits, so the prominence of the ridges increases.
Public-domain text, read in full here on John Shaqi.
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