Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
His best piece of purely morphological work was his theory of the
metamorphosis of plants. Stripped of its vaguer elements, and of the
crude attempt to explain differences in the character of plant organs
by differences in the degree of "refinement" of the sap supplied to
them, the theory is that stem-leaves, sepals, petals, and stamens are
all identical members or appendages. These appendages differ from one
another only in shape and in degree of expansion, stem-leaves being
expanded, sepals contracted, petals expanded, and so on alternately.
It is equally correct to call a stamen a contracted petal, and a petal
an expanded stamen, for no one of the organs is the type of the
others, but all equally are varieties of a single abstract
plant-appendage.
What Goethe considered he had proved for the appendages of plants he
extended to all living things. Every living thing is a complex of
living independent beings, which "der Idee, der Anlage nach," are the
same, but in appearance may be the same or similar, different or
unlike.[80] Not only is there a primordial animal and a primordial
plant, schematic forms to which all separate species are referable,
but the parts of each are themselves units, which "der Idee nach," are
identical _inter se_. This fantasy can hardly be taken seriously as a
scientific theory; it seems, however, to have been what guided Goethe
in his "discovery" of the vertebral nature of the skull. Just as the
fore limb can be homologised with the hind limb, so, reasoning by
analogy, the skull should be capable of being homologised with the
vertebrae. To what ludicrous extremes this doctrine of the repetition
of parts within the organism was pushed we shall see when we consider
the theories of the German transcendentalists of the early nineteenth
century.
Though Goethe's morphological views were lacking in definiteness he
hit upon one or two ideas which proved useful. Thus he enunciated the
"law of balance" long before Etienne Geoffroy St Hilaire, the law
"that to no part can anything be added, without something being taken
away from another part, and _vice versa_."[81] He saw, too, what a help
to the interpretation of adult structure the study of the embryo would
be, for many bones which are fused in the adult are separate in the
embryo.[82] This also was a point to which the later transcendentalists
gave considerable attention.
Public-domain text, read in full here on John Shaqi.
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