Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Prof. Lignier[175] has recently described some large megasporangia
from the Westphalian Coalfield of Ostrau in Austrian Silesia which he
made the type of a new genus _Mittagia_, after Herr Mittag, Director
of Mines. Two sporangia, between 2 and 3 mm. in diameter, were found
in close association as though belonging to a single sorus; one was
empty and the other contained four megaspores. The structure of the
thick wall of the sporangia is very similar to that of the testa
of _Lagenostoma Lomaxi_, but it apparently split into two valves.
Lignier refers the new type, _Mittagia seminiformis_, to some
unknown Palaeozoic group of heterosporous Filicineae, possibly the
ancestral stock of the Pteridosperms, and he thinks it probable that
the sporangia resembled seeds in their facilities for dispersal. In
the structure of the sporangial wall _Mittagia_ also resembles the
sporocarp of _Pilularia_.
_Seeds presented as impressions, without internal structure,
superficially resembling Lagenostoma._
_Lagenospermum_ Nathorst.
This generic name is adopted for seeds represented by casts or
impressions agreeing in external features with _Lagenostoma_ but
which on the available evidence cannot be confidently assigned to
that genus[176]. Two types of seed were described by Arber[177]
from the Lower Coal Measures of Scotland as _Lagenostoma Kidstoni_
and _L. Sinclairi_: the former has been removed by Oliver[178] to
_Physostoma_ and both are included by Arber[179] in a recent paper in
the genus _Radiospermum_. This new generic term is proposed by Arber
for a number of small sub-cylindrical seeds founded on impressions
including ‘small seeds which, when the structure is preserved,
are known as _Lagenostoma_, _Physostoma_, and _Conostoma_.’ The
question of nomenclature is invariably raised by cases in which
impressions resemble in their superficial characters genera founded on
anatomical characters: the seeds originally referred to _Lagenostoma
Sinclairi_ afford a good example of this difficulty. Nathorst has
recently proposed the generic name _Lagenospermum_ as preferable
to _Lagenostoma_ and _Radiospermum_ in the case of _Lagenostoma
Sinclairi_ and similar seeds which afford no proof of the possession
of such morphological characters as would justify their inclusion in
the genus _Lagenostoma_ but which may be examples of that genus. As
Nathorst points out, the adoption of _Radiospermum_ for _L. Sinclairi_
is inadvisable on the ground that it is also applied to seeds of a
different type. The type-species of _Lagenospermum_ is _L. Sinclairi_
and Nathorst describes additional species from Lower Carboniferous
rocks in Spitzbergen.
_Lagenospermum Sinclairi_ (Arber ex Kidston +MS.+).
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