In Weltner's catalogue of the freshwater sponges (1895) seventy-six
recent species of Spongillidæ (excluding _Lubosmirskia_) are enumerated,
and the number now known is well over a hundred. In India we have
twenty-nine species, subspecies, and varieties, while from the whole of
Europe only about a dozen are known. In the neighbourhood of Calcutta
nine species, representing three genera and a subgenus, have been found;
all of them occur in the Museum tank. The only other region of similar
extent that can compare with India as regards the richness of its
freshwater sponge fauna is that of the Amazon, from which about twenty
species are known. From the whole of North America, which has probably
been better explored than any other continent so far as Spongillidæ are
concerned, only twenty-seven or twenty-eight species have been recorded.
The Indian species fall into seven genera, one of which (_Spongilla_)
consists of three subgenera. With one exception (that of
_Pectispongilla_, which has only been found in Southern India) these
genera have a wide distribution over the earth's surface, and this is
also the case as regards the subgenera of Spongilla. Four genera
(_Heteromeyenia_, _Acalle_, _Parmula_, and _Uruguaya_) that have not yet
been found in India are known to exist elsewhere.
Five of the Indian species are known to occur in Europe, viz.,
_Spongilla lacustris_, _S. crateriformis_, _S. carteri_, _S. fragilis_,
_Trochospongilla pennsylvanica_; while _Ephydatia meyeni_ is
intermediate between the two commonest representatives of its genus in
the Holarctic Zone, _Ephydatia fluviatilis_ and _E. mülleri_. Of the
species that occur both in India and in Europe, two (_Spongilla
lacustris_ and _S. fragilis_) are found in this country in forms
sufficiently distinct to be regarded as subspecies or local races.
Perhaps this course should also be taken as regards the Indian forms of
_S. carteri_, of which, however, the commonest of the Indian races would
be the typical one; but _S. crateriformis_ and _T. pennsylvanica_ seem
to preserve their specific characters free from modification, whether
they are found in Europe, Asia, or America.
The freshwater sponges of Africa have been comparatively little studied,
but two Indian species have been discovered, _S. bombayensis_ in Natal
and _S. alba_ var. _cerebellata_ in Egypt. Several of the species from
the Malabar Zone are, moreover, closely allied to African forms (p. 11).
FOSSIL SPONGILLIDÆ.
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