The Spongillidæ are an ancient family. Young described a species
(_Spongilla purbeckensis_) from the Upper Jurassic of Dorset (Geol. Mag.
London (new series) v, p. 220 (1878)), while spicules, assigned by
Ehrenberg to various genera but actually those of _Spongilla lacustris_
or allied forms, have been found in the Miocene of Bohemia (see
Ehrenberg's 'Atlas für Micro-Geologie,' pl. xi (Leipzig, 1854), and
Traxler in Földt. Közl., Budapest, 1895, p. 211). _Ephydatia_ is also
known in a fossil condition, but is probably less ancient than
_Spongilla_.
Ehrenberg found many sponge spicules in earth from various parts of the
Indian Empire (including Baluchistan, Mangalore, Calcutta, the Nicobars
and Nepal) and elsewhere, and it might be possible to guess at the
identity of some of the more conspicuous species figured in his 'Atlas.'
The identification of sponges from isolated spicules is, however, always
a matter of doubt, and in some cases Ehrenberg probably assigned
spicules belonging to entirely different families or even orders to the
same genus, while he frequently attributed the different spicules of the
same species to different genera. Among his fossil (or supposed fossil)
genera that may be assigned to the Spongillidæ wholly or in part are
_Aphidiscus_, _Spongolithis_, _Lithastericus_ and _Lithosphæridium_,
many of the species of these "genera" certainly belonging to _Spongilla_
and _Ephydatia_.
ORIENTAL SPONGILLIDÆ NOT YET FOUND IN INDIA.
Few freshwater sponges that have not been found in India are as yet
known from the Oriental Region, and there is positive as well as
negative evidence that Spongillidæ are less abundant in Malaysia than in
this country. The following list includes the names of those that have
been found, with notes regarding each species. It is quite possible that
any one of them may be found at any time within the geographical
boundaries laid down for this 'Fauna.' I have examined types or co-types
in all cases except that of _Ephydatia fortis_, Weltner.
I. _Spongilla_ (_Euspongilla_) _microsclerifera_*, Annandale
(Philippines). P. U.S. Mus. xxxvii, p. 131 (1909).
This sponge is closely related to _S. lacustris_, but apparently does
not produce branches. It is remarkable for the enormous number of
microscleres in its parenchyma.
II. _S._ (_Euspongilla_) _philippinensis_*, Annandale (Philippines). P.
U.S. Mus. xxxvi, p. 629 (1909).
Related to _S. alba_ and still more closely to _S. sceptrioides_ of
Australia. From the former it is readily distinguished by having
minutely spined megascleres, green corpuscles, slender gemmule-spicules
with short spines and no free microscleres.
III. _S._ (? _Euspongilla_) _yunnanensis_*, Annandale (W. China). Rec.
Ind. Mus. v, p. 197 (1910).
Apparently allied to _S. philippinensis_ but with smooth
skeleton-spicules and a more delicate skeleton.
IV. _S._ (_Stratospongilla_) _sinensis_*, Annandale (Foochow, China). P.
U.S. Mus. xxxviii, p. 183 (1910).
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