II. Some or all of the microscleres birotulate.
(Birotulate microscleres of one kind only.)
A. Microscleres of the gemmule birotulate, the
rotules with serrated or strongly sinuous
edges; parenchyma spicules usually absent,
never of complicated structure EPHYDATIA, p. 108.
B. Microscleres of the gemmule as in
_Ephydatia_; microscleres of the parenchyma
consisting of numerous shafts
meeting in different planes in a central
nodule DOSILIA, p. 110.
C. Microscleres as in _Ephydatia_ except
that the rotulæ of the gemmule-spicules
have smooth edges TROCHOSPONGILLA, p. 113.
D. Microscleres of the gemmule without a
trace of rotules, those of the parenchyma
birotulate CORVOSPONGILLA, nov., p. 122.
III. Microscleres of the gemmule with a well-developed
basal rotule and a vertical shaft
ending above in a mere knob. TUBELLA, p. 120.
The most distinct genus of Spongillidæ not yet found in India is
_Heteromeyenia_, Potts. It is easily distinguished from all others by
the fact that the birotulate spicules of the gemmule are of two quite
distinct kinds, which occur together on every mature gemmule.
_Heteromeyenia_ is represented by several American species, one of which
has been found in Europe. _Acalle_, J. E. Gray, which is represented by
a single South American species (_Spongilla recurvata_, Bowerbank), is
related to _Heteromeyenia_ but has one kind of gemmule-spicule
tubelliform, the other birotulate. Probably _Uraguaya_, Carter, should
be regarded as a subgenus of _Trochospongilla_ with an unusually solid
skeleton; it is peculiar to S. America. _Parmula_, Carter (=_Drulia_,
Gray) includes South American forms allied to _Tubella_, but with the
shaft of the gemmule-spicule degenerate and consisting of a mere
projection in the centre of a shield-like body, which represents the
lower rotule. The status of _Potamolepis_, Marshall, originally
described from the Lake of Galilee, is very doubtful; possibly some or
all of its species belong to the subgenus of _Spongilla_ here called
_Stratospongilla_ (p. 100); but they are stated never to produce
gemmules. The same is the case as regards _Pachydictyum_, Weltner, which
consists of a single species from Celebes.
The sponges from Lake Baikal assigned by Weltner (Arch. Naturg. lxi (i)
p. 131) to the subfamily Lubomirskinæ are of doubtful position and need
not be considered here; while _Lessepsia_, Keller, from one of the salt
lakes on the Suez Canal, certainly does not belong to the family,
although it is assigned to it by von Lendenfeld (Mon. Horny Sponges, p.
904 (1889)) and subsequently by Minchin (Porifera, p. 152, in
Lankester's Treatise on Zoology, part ii (1900)).
Genus 1. SPONGILLA, _Lamarck_ (Carter _emend._).
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