Grasses : $b A handbook for use in the field and laboratoryWard, H. Marshall (Harry Marshall)
Science
Grasses : $b A handbook for use in the field and laboratory
Ward, H. Marshall (Harry Marshall)
Grasses
Each vascular bundle has its own sclerenchyma sheath, and very often
the stronger veins are accentuated owing to the vascular bundle
having a girder-like band of sclerenchyma running conjointly with
its sheath and joining the latter above and below--or below only--to
the epidermis (Figs. 24 and 25). In many cases these lower girders
spread out laterally below--fan-shaped in section--and nearly join the
neighbouring girders.
In other cases the strands of sclerenchymatous supporting tissue do not
join the bundles, but run parallel to them, above or below, as separate
strands just beneath the epidermis.
Finally, these strands may separate from the bundles, and fuse below
into a continuous layer under the epidermis; this occurs especially
in leaves of xerophytes where the cuticle is well developed--e.g. in
varieties of _Festuca ovina_ (Fig. 18), _Aira flexuosa_ (Fig. 28).
The distribution of the strands of isolated sclerenchyma affords
good characters. While there are none in _Mibora_, we find one large
strand at the ridge of the keel and one at each margin, in addition to
smaller ones subtending each vascular bundle, in _Avena pubescens_,
_Sesleria_, _Poa annua_ (Fig. 21), _P. bulbosa_, _P. compressa_ and
_Dactylis glomerata_. In _Festuca ovina_, _F. rubra_, _F. heterophylla_
(Figs. 18, 27) there are groups more or less pronounced at the keel and
margins, or even a continuous band below, but none above the bundles.
[Illustration: Fig. 26. Transverse section of leaf of _Nardus stricta_
(× about 50). The upper surface is represented by the four grooves and
five ridges, each of the former with traces of motor-cells at its base.
The deep shaded portions are sclerenchyma, strong girders of which
join the vascular bundle of each ridge to the lower surface. This type
is obviously derived from that in Fig. 19, and may be regarded as a
permanently rolled leaf.]
[Illustration: Fig. 27. Transverse section of leaf of _Festuca ovina_,
var. _duriuscula_ (× about 50), the type of a permanently folded leaf.
Seven ridges and six intervening grooves are seen: each of the latter
with traces of motor-cells below. In each ridge is an isolated vascular
bundle, and a narrow sclerenchyma band below.]
Many grasses have an isolated band above and below each primary bundle
only--e.g. _Panicum_, _Cynodon_--or above and below each of the other
bundles as well--e.g. _Spartina_, _Arundo_, _Polypogon_, _Agrostis
alba_, _Aira cæspitosa_ (Fig. 23), _Holcus lanatus_, _Glyceria
aquatica_, _G. fluitans_, _Digraphis_, _Elymus_ (Fig. 25), _Agropyrum_
(Fig. 24), _Brachypodium_, _Nardus_ (Fig. 26). In _Psamma arenaria_ the
lower bands join into a continuous layer.
In the following there is a band like a girder above and below each
bundle, and contiguous with it, joining it to the epidermis above and
below--_Leersia_, _Phleum pratense_, _Calamagrostis Epigeios_, _Bromus
erectus_, &c.
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