Handbook of Medical EntomologyRiley, William A. (William Albert)
Science
Handbook of Medical Entomology
Riley, William A. (William Albert)
Insect pests; Insects as carriers of disease; Medical parasitology
The relation of insects to man as simple parasites has long been
studied, and until very recent years the bulk of the literature of
medical entomology referred to this phase of the subject. This is now
completely overshadowed by the fact that so many of these parasitic
forms are more than simple parasites, they are transmitters of other
microscopic parasites which are pathogenic to man. Yet the importance of
insects as parasites still remains and must be considered in a
discussion of the relation of insects to the health of man. In taking up
the subject we shall first consider some general features of the
phenomenon of animal parasitism.
Parasitism is an adaptation which has originated very often among living
organisms and in widely separated groups. It would seem simple to define
what is meant by a "parasite" but, in reality, the term is not easily
limited. It is often stated that a parasite is "An organism which lives
at the expense of another," but this definition is applicable to a
predatory species or, in its broadest sense, to all organisms. For our
purpose we may say with Braun: "A parasite is an organism which, for the
purpose of obtaining food, takes up its abode, temporarily or
permanently, on or within another living organism".
Thus, parasitism is a phase of the broad biological phenomenon of
_symbiosis_, or living together of organisms. It is distinguished from
_mutualism_, or symbiosis in the narrow sense, by the fact that only one
party to the arrangement obtains any advantage, while the other is to a
greater or less extent injured.
Of parasites we may distinguish on the basis of their location on or in
the host, _ecto-parasites_, which live outside of the body; and
_endo-parasites_, which live within the body. On account of their method
of breathing the parasitic arthropods belong almost exclusively to the
first of these groups.
On the basis of relation to their host, we find _temporary parasites_,
those which seek the host only occasionally, to obtain food; and the
_stationary_ or _permanent_ parasites which, at least during certain
stages, do not leave their host.
_Facultative parasites_ are forms which are not normally parasitic, but
which, when accidentally ingested, or otherwise brought into the body,
are able to exist for a greater or less period of time in their unusual
environment. These are generally called in the medical literature
"pseudoparasites" but the term is an unfortunate one.
We shall now take up the different groups of arthropods, discussing the
more important of the parasitic forms attacking man. The systematic
relationship of these forms, and key for determining important species
will be found in Chapter XII.
ACARINA OR MITES
The ACARINA, or _mites_, form a fairly natural group of arachnids,
characterized, in general, by a sac-like, unsegmented body which is
generally fused with the cephalothorax. The mouth-parts have been united
to form a beak or rostrum.
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