Goethe in 1790 revived Wolff's theory of the flower, without suspicion
that he had been anticipated. It is only our ignorance, he said, when
the fact came to his knowledge, that ever deludes us into believing
that we have put forth an original view. As soon as he realised the
true state of the case, he spared no pains to do Wolff full justice.
The aim of Goethe's _Metamorphoses of Plants_ was to determine the
Idea or theoretical conception of the plant, and also to trace the
modifications which the Idea undergoes in nature. These two inquiries
constituted what he called the Morphology of the plant, a useful,
nay, indispensable term, which is still in daily use. He thought
that he could discover in the endless variety of the organs of
the flowering plant one structure repeated again and again, which
gradually attained, as by the steps of a ladder, what he called the
crowning purpose of nature—viz., the sexual propagation of the race.
This fundamental structure was the leaf. The proposition that all the
parts of the flower are modifications of the leaf he defended by three
main arguments—viz., (1) the structural similarity of seed-leaves,
foliage-leaves, bracts, and floral organs; (2) the existence of
transitions between leaves of different kinds; and (3) the occasional
_retrogression_, as he called it, of specially modified parts to a
more primitive condition. These lines of argument were illustrated by
many well-chosen examples, the result of long and patient observation.
Goethe did not, however, fortify his position by the likeness of
developing floral organs to developing foliage-leaves, which had been
Wolff's starting-point. He arrived independently at Wolff's opinion
that the conversion of foliage-leaves into floral organs is due to
diminished nutrition.
Linnæus's exposition of the nature of the flower had been read
attentively by Goethe, who must have remarked that the conversion of
organs to new uses was there described as a _metamorphosis_. That
word had been, long before the time of Linnæus, appropriated to a
particular kind of change—viz., an apparently sudden change occurring
in the life-history of one and the same animal. It was therefore unlucky
that Goethe should have been led by the example of Linnæus to employ the
word in the general sense of adaptation to new purposes. He did not,
however, expressly compare flower-production with the transformation of
an insect, as Linnæus had done.
The reception of Goethe's _Metamorphosen der Pflanzen_ was at first
cold, but the doctrine which it enforced gradually won the attention of
botanists, and by 1830 he was able to show that it had been accepted by
many good judges.
Public-domain text, read in full here on John Shaqi.
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