The fathers of botany neglected everything else in order to concentrate
their attention upon the flowering plants, from which very nearly
all useful vegetable products were derived. The lack of adequate
microscopes rendered it almost impossible to investigate the structure
and life-history of ferns, mosses, fungi, and algæ until the nineteenth
century. As late as the time of Linnæus it was possible to maintain
that they developed spontaneously, though the great naturalist himself
called them Cryptogamia, thereby expressing his conviction that they
reproduce their kind like other plants, but in a way so far not
understood. Gaertner, a contemporary of Linnæus, pointed out one
important respect in which the spores of cryptogams differ from the
seeds of flowering plants, viz. that they contain no embryo.
_Ferns._—Even before the age of Linnæus it was known that little
ferns spring up around the old ones, and that a fine dust can be shaken
from the brown patches on the back of ripe fern-leaves. The dust was
reputed to be the seed of the fern, and in an age which believed in
magic the invisibility of fern-seed made it easy to suppose that the
possessor of fern-seed would become invisible also. When the microscope
began to be applied to minute natural objects, the brown patches of the
fern-leaf were closely examined. William Cole of Bristol (1669),
Malpighi, Grew, Swammerdam, Leeuwenhoek, and others, found the stalked
capsules (sporangia), their elastic ring and the minute bodies (spores)
lodged within them; it seemed obvious to call the capsules ovaries and
the spores seeds. Some time in the latter part of the seventeenth
century Robert Morison, professor of botany at Oxford, who died in 1683,
sowed spores of the harts-tongue fern, and next year got an abundant
crop of prothalli, which he took to be the cotyledons. A little later,
when it had been proved that flowering plants possess male and female
organs, diligent search was made for the stamens and pistils of ferns
and mosses, which of course could not be found, though identifications,
sometimes based upon a real analogy, were continually announced. Late
in the eighteenth century one John Lindsay, a surgeon in Jamaica, who
was blest with leisure and a good microscope, repeated the experiment
of Morison, which seems to have been almost forgotten. Having remarked
that after the rains young ferns sprang up in shady places where the
earth had been disturbed. it occurred to him to mix the fine brown
dust from the back of a fern-leaf with mould, sow the mixture in a
flower-pot, and watch daily to see what might come up. About the
twelfth day small green protrusions were observed, which enlarged,
sent down roots, and formed bilobed scales, out of which young ferns
ultimately grew. In 1789 Sir Joseph Banks, who was reputed to be the
best English botanist of the day, asked Lindsay's help in sending West
Indian ferns to Europe. Lindsay replied that it would be easier to send
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