reflexed hairs allow small flies to creep in, but effectually prevent
their escape until they have fertilised the pistils, when the hairs
relax. These are only specimens of a multitude of adaptations which
fill the book.
Sprengel insists upon the study of flowers under natural conditions;
he could never have made out by the examination of plucked flowers
how Nigella is fertilised. Flies with attached pollen-masses, which
he found in spiders' nests, gave him the hint as to the way in which
the fertilisation of orchids is effected. Definite questions must
be put if observation is to be profitable. What is the use of honey
to the plant—of this coloured spot—of these hairs? He notes the
peculiarities of wind-fertilised and insect-fertilised flowers, the
relative abundance of the pollen, the form of the stigma, the presence
or absence of honey, the size, colour, and scent of the corolla. Here
is a pretty illustration from his pages. Pluck a branch of hazel,
aspen, or alder, with unexpanded catkins, and also one from the male
sallow; place them in water, and keep them in a sunny window until
the anthers are ripe. A vigorous puff will then discharge a cloud
of pollen from the _wind-fertilized_ catkins, but none from the
_insect-fertilised_ catkin of the sallow. What Linnæus said about
the flowers of trees appearing before the leaves, in order that the
pollen may more easily reach the stigmas, holds good, Sprengel remarks,
only of wind-fertilised trees. The lime, which is insect-fertilised,
flowers in the height of summer, when all the branches are crowded with
leaves.
Sprengel left it to later biologists to complete his discovery. "That
wonderfully accurate observer, Sprengel," says Darwin,[32] "who first
showed how important a part insects play in the fertilisation of
flowers, called his book _The Secret of Nature Displayed_; yet he
only occasionally saw that the object for which so many curious and
beautiful adaptations have been acquired, was the cross-fertilisation
of distinct plants; and he knew nothing of the benefits which the
offspring thus receive in growth, vigour, and fertility." Not even
Cuvier and the Rise of Palæontology.
If this historical sketch had been prepared within a few years of the
death of Cuvier, it would no doubt have held him up as the greatest
of zoologists and comparative anatomists. Nor would it have been hard
to find reasons for such a verdict. His _Règne Animal_ extended and
corrected the zoological system of Linnæus; his comparative anatomy,
and especially his comparative osteology, were far ampler and more
exact than anything that had been attempted before. It would not have
been forgotten, moreover, that he was the practical founder of the new
science of palæontology.
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