Life Movements in Plants, Volume IBose, Jagadis Chandra
Science
Life Movements in Plants, Volume I
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
I may here say a few words of the manner in which the period of
transmission can be found from the record given by my Resonant
Recorder, fully described in my previous paper. The writer attached to
the recording lever of this instrument is maintained by electromagnetic
means in a state of to-and-fro vibration. The record thus consists
of a series of dots made by the tapping writer, which is tuned to
vibrate at a definite rate, say, 10 times per second. In a particular
case whose record is given in Curve 1 (Fig. 46), indirect stimulus
of electric shock was applied at a distance of 15 mm. from the
responding pulvinus. There are 15 intervening dots between the
moment of application of stimulus and the beginning of response; the
time-interval is therefore 1.5 seconds. The latent period of the
motile pulvinus is obtained from a record of direct stimulation; the
average value of this in summer is 0.1 second. Hence the true period
of transmission is 1.4 seconds for a distance of 15 mm. The velocity
determined in this particular case is therefore 10.7 mm. per second.
Precaution has to be taken against another source of disturbance,
namely, the excitation caused by the sudden commencement or the
cessation of the constant current. I have shown elsewhere[O] that the
sudden initiation or cessation of the current induces an excitatory
reaction in the plant-tissue similar to that in the animal tissue. This
difficulty is removed by the introduction of a sliding potentiometer,
which allows the applied electromotive force to be gradually increased
from zero to the maximum or decreased from the maximum to zero.
[O] BOSE--‘Plant Response’ (1906); ‘Irritability of Plants’ (1913).
[Illustration: FIG. 45.--Complete apparatus for investigation of
the variation of conducting power in _Mimosa_. A, storage cell; S,
potentiometer slide, which, by alternate movement to right or left,
continuously increases or decreases the applied E. M. F.; K, switch key
for putting current “on” and “off” without variation of resistance; E,
E′, electrodes of induction coil for stimulation; C, choking coil; G,
micro-ammeter.]
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