Life Movements in Plants, Volume IIBose, Jagadis Chandra
Science
Life Movements in Plants, Volume II
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
Before entering into the fuller consideration of the subject, it will be
helpful to form some mental picture of the phenomena of excitation,
however inadequate it may be. The excitation is admitted to be due to
the molecular upset induced by the shock of stimulus[17]; the increased
excitation results from increasing molecular upset brought on by
enhanced stimulus. The condition of molecular upset or excitation may be
detected from the record of any one of the several concomitant changes,
such as the change of form, (contraction or expansion) or change of
electric condition (galvanometric negativity or positivity). These means
of investigation are not in principle different from those we employ in
the detection of molecular distortion in inorganic matter under
increasing intensities of an external force.
[17] I shall use the term _stimulus_ in preference to _stimulation_,
for the latter is often taken in the sense of the resulting
excitation.
THE CHARACTERISTIC CURVE.
Thus the molecular upset and rearrangement, in a magnetic substance
under increasing magnetising force are inferred from the curve obtained
by means of appropriate magnetometric or galvanometric methods. I
reproduce the characteristic curve of iron (Fig. 131) in which the
abscissa represents increasing magnetising force, and the ordinate, the
induced magnetisation. This characteristic curve, giving the relation of
cause and effect, will be found to be highly suggestive as regards the
similar characteristic curve which gives the relation between increasing
stimulus and the resulting enhanced tropic effect in vegetable tissues.
The parallelism will be found to be very striking.
Inspection of figure 131 shows that, broadly speaking, the curve of
magnetisation may be divided into four parts. In the first part, under
feeble magnetic force, the slope of the curve is very small; later, in
the second part, as the force increases, the curve becomes very steep;
in the third part the slope of the curve remains fairly constant; and
finally in the fourth part, the curve rounds off and the rate of ascent
again becomes very small. The _susceptibility_ for induced magnetisation
is thus very feeble at the beginning; under increasing force, in the
second stage, the susceptibility becomes greatly enhanced; in the third
stage, the susceptibility remains approximately constant; and in the
fourth stage it becomes diminished. We shall presently find that the
susceptibility for excitation also undergoes a similar variation at the
four different stages of stimulation.
CHARACTERISTICS OF SIMPLE PHOTOTROPIC CURVE.
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