Links With the Past in the Plant WorldSeward, A. C. (Albert Charles)
Science
Links With the Past in the Plant World
Seward, A. C. (Albert Charles)
Paleobotany -- Great Britain; Plants -- Great Britain
approaching the island everyone was satisfied that the columnar objects
were trees, 'except our Philosophers, who still maintained they were
basaltes.' The leaves are short, about half an inch long, laterally
compressed and slightly spreading and sickle-shaped--sometimes shorter
and broader and overlapping--arranged in crowded spirals. The scales
of the broadly oval cones are single-seeded, but differ from those
of _Araucaria imbricata_ in having the seed exposed on the surface
and in the greater breadth and thinner borders of the scales. In both
Araucaria and Agathis the nature of the seed-scales constitutes a
distinguishing feature. The leaves of _Araucaria imbricata_ differ in
form from those of other Conifers. The foliage shoots of _Araucaria
excelsa_ and other species, _e.g._ the very closely allied _A. Cookii_
of the New Hebrides and New Caledonia, though not unlike the branches
of a Japanese Conifer (_Cryptomeria japonica_), often cultivated in
England, afford fairly trustworthy characters for identification
purposes.
The minute structure of the wood of both Araucaria and Agathis
constitutes an important distinguishing feature and enables us to
recognise on microscopical examination even a fragment of wood of
either of these genera. The small elongated cells or water-conducting
elements of the wood of the Araucarieae are characterised by one or
two, and occasionally as many as three or four, contiguous rows
of pits on their radial walls, and these appear in surface view as
flattened circles or polygonal areas.
These details have been mentioned in order to show that Araucaria and
Agathis are sufficiently distinct in many respects from other Conifers
to render their identification in a fossil state comparatively easy, at
least much easier than the recognition of the majority of the members
of the Coniferae. It would be going too far to state definitely that
Araucarieae, as defined by reference to existing species, existed
during the Palaeozoic period; on the other hand it would seem in a
high degree probable that the vegetation of the Coal age and of the
succeeding Permian period included trees in which certain Araucarian
characters were clearly foreshadowed. The name Araucarioxylon was
formerly applied to petrified wood, obtained from Palaeozoic as well
as from later formations, which agrees anatomically with that of
Araucaria and Agathis. It has been shown in recent years that much of
the Palaeozoic wood of this type of structure belongs to the extinct
genus Cordaites, a tree which played a prominent part in the earlier
floras. Cordaites affords a good example of a generalised type: in its
wood-structure it resembles very closely the existing Araucarieae;
its long strap-like leaves are not unlike those of some species of
Agathis; its male flowers have often been compared with those of the
Maiden Hair tree, _Ginkgo biloba_, and certain anatomical features form
connecting links between this Palaeozoic genus and the Cycads.
Public-domain text, read in full here on John Shaqi.
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