The male and female flowers are borne on separate trees; the male consists of a central axis giving off slender branches, each of which ends in a small terminal knot and two elliptical capsules in which the pollen is produced. The female flowers have a stouter axis which normally produces two seeds at the apex. The seed is encased in a green fleshy substance and, as in the fruit of a cherry or plum, the kernel is protected by a hard woody shell. In the form of the leaves and in the structure of the flowers Ginkgo presents features which clearly distinguish it from the Conifers, the class in which, until recently, it was included. In 1896 the Japanese botanist Hirase made the important discovery that the male reproductive cells of Ginkgo are large motile bodies provided with a spirally coiled band of minute cilia--delicate hairs which by their rapid lashing-movement propel the cell through water. In all Flowering Plants and in Conifers the male reproductive cells have no independent means of locomotion; they are carried to the female cell by the formation of a slender tube--the pollen-tube--produced by the pollen-grain. In the Ferns, Lycopods and Horsetails--in fact in all members of the Pteridophyta--as also in the Mosses and Liverworts as well as in many of the still lower plants, the male cells swim to the egg by the lashing of cilia like those on the male cells of Ginkgo. This difference in regard to the nature of the male cells was considered to be a fundamental distinction between the higher seed-bearing plants and all other groups of the vegetable kingdom. It was, therefore, with no ordinary interest that Hirase's discovery was received, as it broke down a distinction between the two great divisions of the plant-world which had been generally accepted as fundamental; though it is only fair to say that the German botanist Hofmeister, a man of exceptional originality and power of grasping the essential, foresaw the possibility that this arbitrary barrier would eventually be removed. The Ferns and other plants in which the male cells are motile, represent earlier stages in the progress of plant development, when the presence of water was essential for the act of fertilisation, a relic of earlier days when the whole plant-body was fitted for a life in water. As higher types were produced, the plant-machinery became less dependent on an aqueous habitat, and the loss of organs of locomotion in the male cells is an instance of the kind of change accompanying the gradual adaptation to life on land. The idea of the gradual emancipation of plants from a watery environment is expressed in a somewhat extreme form by the author of a book entitled _The Lessons of Evolution_(52), who states that the ocean is the mother of plant-life and that plants formed the army which conquered the land. In Ginkgo we have a type which, though similar in most respects to the Conifers, possesses in its motile reproductive cells a persistent inheritance from the past.