Memoir on the Dodo (Didus ineptus, Linn.)Owen, Richard
History
Memoir on the Dodo (Didus ineptus, Linn.)
Owen, Richard
Dodo
The ribs of the Dodo are as broad, in proportion to their length, as in
Doves, but are relatively longer in proportion to the dorsal region,
encompassing a more capacious thoracic-abdominal cavity. The ribs
of the Vulture are more expanded than in _Didus_, especially where
they afford the extensive attachment to the epipleurals. But I shall
not dwell further on the comparative characters of this part of the
skeleton, as more decisive ones of the affinity of _Didus_ are afforded
by other parts.
In comparing the sternum of the Dodo with that of Doves of flight,
the first well-marked difference is in the adaptive development of
the keel in the last (Pl. III. fig. 2, _Didunculus_), and in the
provision for the concomitantly broader coracoids, the grooves for
which meet and run into each other across the fore part of the bone in
existing _Columbidæ_ (Pl. XII. fig. 2, _b_); consequently the inner
or upper wall of the confluent grooves forms a median prominence (ib.
_e_) at the front margin of the sternum, contrasting with the wide
notch at that part of the bone in the Dodo (Pl. IV. fig. 4). The next
difference, as compared with _Goura_ and most Pigeons, is the absence
of the entolateral processes (Pl. XII. fig. 3, _i_) in the Dodo’s
sternum: but _Didunculus_ singularly exemplifies its nearer affinity
to _Didus_ by a like absence of those processes; only the sternal
margins behind the ectolateral processes (ib. fig. 1, _h_), instead of
converging with a slight convexity to an obtuse apex, as in Pl. VI.,
describe a concavity, through an expansion of the posterior truncate
end of the breast-bone. The sternum of _Didunculus_ may be said to
show one pair of posterior notches (Pl. XII. fig. 1, _f_), that of
other Pigeons two pairs (ib. fig. 3, _f_ _f′_); but the sternum of
_Didus_, which is relatively broader, shows no other trace of the
anterior notch (Pl. VI. _f_) than is afforded by the rounded angle at
which the ectolateral process (_h_) rises from the bone. Although the
costal margin is relatively shorter in Doves of flight than in the
Dodo, again an intermediate condition is manifested by _Didunculus_
as compared with _Goura_, in which latter Dove there are articular
surfaces for three sternal ribs (Pl. XII. fig. 3, _o_ 1, 2, 3),
whilst in _Didunculus_ there are four (ib. fig. 1, _c_). _Didunculus_
also exhibits, more strongly than _Goura_, the obtuse ridges (ib.
fig. 2, _r_) converging like buttresses from the outer wall of the
coracoid groove to the fore part of the keel, where they subside. In
_Didunculus_ there is a pneumatic foramen exterior to the coracoid
groove, corresponding with _p_, fig. 4, Pl. IV., which I do not find
in the sternum of _Goura_; but in the Crown-pigeons the pneumatic
foramina along the middle line of the upper surface of the sternum are
conspicuous; they are confined to the fore part of that surface in
_Didunculus_ (Pl. XII. fig. 1).
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