Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
The first and second experiments had primarily the object of proving
the composition of the hybrid egg cells, while the third and fourth
experiments were to decide that of the pollen cells[40]. As is shown by
the above demonstration the first and second experiments and the third
and fourth experiments should produce precisely the same combinations,
and even in the second year the result should be partially visible in
the form and colour of the artificially fertilised seed. In the first
and third experiments the dominant characters of form and colour, _A_
and _B_, appear in each union, and are also partly constant and partly
in hybrid union with the recessive characters _a_ and _b_, for which
reason they must impress their peculiarity upon the whole of the seeds.
All seeds should therefore appear round and yellow, if the theory be
justified. In the second and fourth experiments, on the other hand,
one union is hybrid in form and in colour, and consequently the seeds
are round and yellow; another is hybrid in form, but constant in the
recessive character of colour, whence the seeds are round and green;
the third is constant in the recessive character of form but hybrid in
colour, consequently the seeds are angular and yellow; the fourth is
constant in both recessive characters, so that the seeds are angular
and green. In both these experiments there were consequently four sorts
of seed to be expected--viz. round and yellow, round and green, angular
and yellow, angular and green.
[40] [To prove, namely, that both were similarly differentiated, and
not one or other only.]
The crop fulfilled these expectations perfectly. There were obtained in
the
1st Experiment, 98 exclusively round yellow seeds;
3rd " 94 " " " "
In the 2nd Experiment, 31 round and yellow, 26 round and green, 27
angular and yellow, 26 angular and green seeds.
In the 4th Experiment, 24 round and yellow, 25 round and green, 22
angular and yellow, 27 angular and green seeds.
A favourable result could now scarcely be doubted; the next generation
must afford the final proof. From the seed sown there resulted for the
first experiment 90 plants, and for the third 87 plants which fruited:
these yielded for the--
1st Exp. 3rd Exp.
20 25 round yellow seeds _AB_
23 19 round yellow and green seeds _ABb_
25 22 round and angular yellow seeds _AaB_
22 21 round and angular green and yellow seeds _AaBb_
In the second and fourth experiments the round and yellow seeds yielded
plants with round and angular yellow and green seeds, _AaBb_.
From the round green seeds plants resulted with round and angular green
seeds, _Aab_.
The angular yellow seeds gave plants with angular yellow and green
seeds, _aBb_.
From the angular green seeds plants were raised which yielded again
only angular and green seeds, _ab_.
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