It was suggested in the last chapter that if a new variation arose as a
sport--as a sudden hereditary variation--and if that variation were,
through resemblance to a different and unpalatable species, to be more
immune to the attacks of enemies than the normal form, it was conceivable
that the newer mimetic sport would become established, and in time perhaps
come to be the only form of the species. We may suppose, for example, that
the A female of _P. polytes_ arose suddenly, and that owing to its likeness
to the presumably distasteful _P. aristolochiae_ it became rapidly more
numerous until in some localities it is the commonest or even the only
form. However, before discussing the establishing of a mimetic form in this
manner we must first deal with certain general results which may be
expected to follow on a process of selection applied to members of a
population presenting variations which are inherited on ordinary Mendelian
lines.
Let us suppose that we are dealing with the inheritance of a character
which depends upon the presence of the genetic factor X; and let us also
suppose that the heterozygous form () is indistinguishable {94} from the
homozygous form () in appearance. In other words the character dependent
upon X exhibits complete dominance. With regard to X then all the members
of our population must belong to one or other of three classes. They may be
homozygous (XX) for X, having received it from both parents, or they may be
heterozygous (Xx) because they have received it from only one parent, or
they may be devoid of X, _i.e._ pure recessives (xx). An interesting
question arises as to the conditions under which a population containing
these three kinds of individuals remains stable. By stability is meant that
with the three kinds mating freely among themselves and being all equally
fertile, there is no tendency for the relative proportions of the three
classes to be disturbed from generation to generation. The question was
looked into some years ago by G. H. Hardy, who shewed that if the mixed
population consist of p XX individuals, 2q Xx individuals and r xx
individuals, the population will be in stable equilibrium with regard to
the relative proportions of these three classes so long as the equation pr
= q^2 is satisfied[48].
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