You allude to the Sicily flora not being peculiar, and this being caused
by its recent elevation (well established) in the main part: you will
find Lyell has put forward this very clearly and well. The Apennines
(which I was somewhere lately reading about) seems a very curious case.
I think Forbes ought to allude a little to Lyell's (22/2. See Letter
19.) work on nearly the same subject as his speculations; not that I
mean that Forbes wishes to take the smallest credit from him or any man
alive; no man, as far as I see, likes so much to give credit to others,
or more soars above the petty craving for self-celebrity.
If you come to any more conclusions about polymorphism, I should be very
glad to hear the result: it is delightful to have many points fermenting
in one's brain, and your letters and conclusions always give one plenty
of this same fermentation. I wish I could even make any return for all
your facts, views, and suggestions.
LETTER 23. TO J.D. HOOKER.
(23/1. The following extract gives the germ of what developed into an
interesting discussion in the "Origin" (Edition I., page 147). Darwin
wrote, "I suspect also that some cases of compensation which have been
advanced and likewise some other facts, may be merged under a more
general principle: namely, that natural selection is continually trying
to economise in every part of the organism." He speaks of the general
belief of botanists in compensation, but does not quote any instances.)
[September 1846].
Have you ever thought of G. St. Hilaire's "loi de balancement" (23/2.
According to Darwin ("Variation of Animals and Plants," 2nd edition,
II., page 335) the law of balancement was propounded by Goethe and
Geoffroy Saint-Hilaire (1772-1844) nearly at the same time, but he gives
no reference to the works of these authors. It appears, however, from
his son Isidore's "Vie, Travaux etc., d'Etienne Geoffroy Saint-Hilaire,"
Paris 1847, page 214, that the law was given in his "Philosophie
Anatomique," of which the first part was published in 1818. Darwin
(ibid.) gives some instances of the law holding good in plants.), as
applied to plants? I am well aware that some zoologists quite reject it,
but it certainly appears to me that it often holds good with animals.
You are no doubt aware of the kind of facts I refer to, such as
great development of canines in the carnivora apparently causing
a diminution--a compensation or balancement--in the small size of
premolars, etc. I have incidentally noticed some analogous remarks on
plants, but have never seen it discussed by botanists. Can you think
of cases in any one species in genus, or genus in family, with certain
parts extra developed, and some adjoining parts reduced? In varieties
of the same species double flowers and large fruits seem something of
this--want of pollen and of seeds balancing with the increased number
of petals and development of fruit. I hope we shall see you here this
autumn.
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