Fleischmann further asserts that Haeckel’s “fundamental biogenetic law”
has utterly collapsed. “Recapitulation” does not occur. Selenka’s figures
of ovum-segmentation show that there are specific differences in the
individual groups. The origin and development of the blastoderm or
germinal disc has nothing to do with recapitulation of the phylogeny. It
is not the case that the embryos of higher vertebrates are
indistinguishable from one another. Even the egg-cell has a specific
character, and is totally different from any unicellular organism at the
Protistan level. The much-cited “gill-clefts” of higher vertebrates in the
embryonic stage are not persistent reminiscences of earlier lower stages;
they are rudiments or primordia shared by all vertebrates, and developing
differently at the different levels; (thus in fishes they become breathing
organs, and in the higher vertebrates they become in part associated with
the organs of hearing, or in part disappear again).
Though Fleischmann’s vigorous protest against over-hastiness in
construction and over-confidence on the part of the adherents of the
doctrine of descent is very interesting, and may often be justified in
detail, it is difficult to resist the impression that the wheat has been
rejected with the chaff.(23)
Even a layman may raise the following objections: Admitting that the great
groups of forms cannot be traced back to one another, the palæontological
record still proves, though it may be only in general outline, that within
each phylum there has been a gradual succession and ascent of forms. How
is the origin of what is new to be accounted for? Without doing violence
to our thinking, without a sort of intellectual autonomy, we cannot rest
content with the mere fact that new elements occur. So, in spite of all
“difficulties,” the assumption of _an actual descent_ quietly forces
itself upon us as the only satisfactory clue. And the fact, which
Fleischmann does not discuss, that even at present we may observe the
establishment of what are at least new breeds, impels us to accept an
analogous origin of new species. Even if the biogenetic law really “finds
its chief confirmation in its exceptions,” even if we cannot speak of a
strict recapitulation of earlier stages of evolution, there are
indisputable facts which are most readily interpreted as reminiscences, as
due to affiliation (ideal or hereditary), with ancestral forms. (Note, for
instance, Weismann’s “prediction,” &c.(24)) Even if Archæopteryx and other
intermediate forms cannot be regarded as connecting links in the strict
sense, _i.e._, as being stages in the actual pedigree, yet the occurrence
of reptilian and avian peculiarities side by side in one organism, goes
far to prove the close relationship of the two classes.
Public-domain text, read in full here on John Shaqi.
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