The simplest unicellular forms of life are to be thought of as at the
beginning of evolution; and, since mechanical causes cannot explain their
ascent, it must be assumed that they have an inherent “phylogenetic
potential of development,” which, working epigenetically, results in
ascending evolution. He leaves us to choose between monophyletic and
polyphyletic evolution, but himself inclines towards the latter,
associating with it a rehabilitation of Wigand’s theory of the primitive
cells. If, in the beginning, primitive forms of life arose (probably as
unicellulars) from the not-living, it is not obvious why we need think of
only one so arising, and, if many did so, why they should not have
inherent differences which would at once result in typically different
evolutionary series and groups of forms. But evolution does not go on _ad
libitum_ or _ad infinitum_, for the capacity for differentiation and
transformation gradually diminishes. The organisation passes from a labile
state of equilibrium to an increasingly stable state, and at many points
it may reach a terminus where it comes to a standstill. Man, the dog, the
horse, the cereals, and fruit trees appear to Reinke to have reached their
goal. The preliminary stages he calls “Phylembryos,” because they bear to
the possible outcome of their evolution the same relation that the embryo
does to the perfect individual. Thus, _Phenacodus_ may be regarded as the
Phylembryo of the modern horse. It is quite conceivable that each of our
modern species may have had an independent series of Phylembryos reaching
back to the primitive cells. But the palæontological record, and
especially its synthetic types, lead Reinke rather to assume that instead
of innumerable series, there have been branching genealogical trees, not
one, however, but several.
These views, together or separately, which are characterised chiefly by
the catch-words “polyphyletic descent,” “labile and stable equilibrium,”
and so on, crop up together or separately in the writings of various
evolutionists belonging to the opposition wing. They are usually
associated with a denial of the theory of natural selection, and with
theories of “Orthogenesis,” “Heterogenesis,” and “Epigenesis.”
We shall discuss them later when we are considering the factors in
evolution. But we must first take notice of a work in which the theories
opposed to Darwinian orthodoxy have been most decisively and aggressively
set forth. As far back as 1892 O. Hamann, then a lecturer on zoology in
Göttingen, gathered these together and brought them into the field,
against Haeckel in particular, in his book “Entwicklungslehre und
Darwinismus.”(27)
Public-domain text, read in full here on John Shaqi.
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