From embryology in particular, as elsewhere in general, we read the
“fundamental biogenetic law,” that evolution is from the general to the
special, from the imperfect to the more perfect, from what is still
indefinite and exuberant to the well-defined and precise, but never from
the special to the special. According to Hamann’s hypothesis we must think
of evolution as going on, so to speak, not about the top but about the
bottom. The phyla or groups of forms are great trunks bearing many
branches and twigs, but not giving rise to one another. Still less do the
little side branches of one trunk bear the whole great trunk of another
animal or plant phylum. But they all grow from the same roots among the
primitive forms of life. Unicellulars these must have been, but not like
our “Protists.” They should be thought of as primitive forms having within
themselves the potentialities of the most diverse and widely separate
evolution-series to which they gave rise, as it were, along diverging
fan-like rays.
It would be instructive to follow some naturalist into his own particular
domain, for instance a palæontologist into the detailed facts of
palæontology, or an embryologist into those of embryology, in order to
learn whether these corroborate the assumptions of the Theory of Descent
or not. It is just in relation to these detailed facts that criticisms or
even denials of the theory have been most frequent. Koken, otherwise a
convinced supporter of the theory, inquires in his “Vorwelt,” _apropos_ of
the tortoises, what has become of the genealogical trees that were
scattered abroad in the world as proved facts in the early days of
Darwinism. He asserts, in regard to _Archæopteryx_, the instance which is
always put forward as the intermediate link in the evolution of birds,
that it does not show in any of its characters a fundamental difference
from any of the birds of to-day, and further, that, through convergent
development under similar influences, similar organs and structural
relations result, iterative arrangements which come about quite
independently of descent. He maintains, too, that the principle of the
struggle for existence is rather disproved than corroborated by the
palæontological record.
In embryology, so competent an authority as O. Hertwig—himself a former
pupil of Haeckel’s—has reacted from the “fundamental biogenetic law.” His
theory of the matter is very much that of Hamann which we have already
discussed; development is not so much a recapitulation of finished
ancestral types as the laying down of foundations after the pattern of
generalised simple forms, not yet specialised; and from these foundations
the special organs rise to different levels and grades of differentiation
according to the type.(29) But we must not lose ourselves in details.
Public-domain text, read in full here on John Shaqi.
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