North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
Lacépède (1788:137, Pl. 7) referred to Garden's description in Pennant
only as "The Molle" but on a folded paper chart entitled "Table Méthodique
des Quadrupèdes ovipares," which is inserted after an introduction of 17 pages,
listed _T. mollis_; this name is again listed on another folded chart, entitled
"Synopsis methodica Quadrupedum oviparorum," which is inserted between
pages 618 and 619 under the genus _Testudo_. The illustration (Pl. 7) was
taken from Pennant (Duméril and Bibron, _loc. cit._). The type locality has
been designated "(following Stejneger, 1944) as eastern Florida" by Schmidt
(1953:108).
Bartram failed to use a binomial name with his description of "the great
soft shelled tortoise," which appeared in his _Travels_ (1791:177-179, Pl. 4 and
unnumbered plate between pages 282 and 283) and two editions of a French
translation (1799 and 1801, 1:307); see Harper (1940). Recently, Bartram's
_Travels_ has been placed on the Official Index of Rejected and Invalid Works
in Zoological Nomenclature, Opinion 447 (see Hemming, 1957). Bartram's
description of a soft-shelled turtle has provided the basis for the proposal of
at least three name-combinations. The first was _Testudo_ (_ferox?_) _verrucosa_
proposed in 1795 by Schoepff; it appeared simultaneously in _The Historia
Testudinum_ and in a German translation, _Naturgeschichte der Schildkröten_
(see Mittleman, 1944:245). Stejneger (1944:26) listed the type locality as
eastern Florida. Daudin (1801:74), also referring to Bartram's description
in his _Voyage_ (French translation), proposed the name _Testudo bartrami_;
Harper (_op. cit._:717) restricted the type locality of _T. bartrami_ from "Halfway
pond," east Florida, to southwestern Putnam County between Palatka
and Gainesville, Florida. Rafinesque (1832:64-65), relying on the authenticity
of the illustrations in Bartram's _Travels_ that depict a soft-shelled turtle
having five claws on each of the hind feet, tubercles on the sides of the head
and neck, and ten scales in the middle of the carapace (presumably inaccuracies
or a composite on the part of the artist), referred to Bartram's description as
a new genus, _Mesodeca bartrami_, a name which Boulenger (1889:245, footnote)
referred to as "mythical." Geoffroy (1809a:18-19) considered Bartram's
description the basis for the recognition of a second species of _Chelys_ (binomial
nomenclature not employed), and Duméril and Bibron (_loc. cit._) suggested
that the description was based partly on a "Chelyde Matamata."
The descriptive comments of Bartram are not clearly applicable to _Testudo
ferox_ Schneider; _Trionyx ferox_, however, is the only species of soft-shelled
turtle known to occur in the region of Bartram's observations (east Florida),
and the type locality was restricted to Putnam County, Florida, by Harper.
The name-combinations, _Testudo___ (_ferox?_) _verrucosa_ Schoepff, _Testudo bartrami_
Daudin, and _Mesodeca bartrami_ Rafinesque are junior synonyms of _Testudo
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