On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
Now all this cannot be exactly calculated in any given case, and it will be
better, instead of haggling about individual cases concerning which we can
never judge with certainty, to take the position adopted in the text and
say: Since the utility of the initial stages _must_ be assumed unless we
are to renounce forever the explanation of adaptation, let us then take it
for granted. No contradiction of facts is involved in this assumption; in
fact, even individual variations exist whose eventual utility can be
demonstrated, for example, the invisible differences enabling Europeans of
certain constitutions to resist the attacks of tropical malarial
fevers,--or the differences of structure, likewise not directly visible,
which enable palms from the summits of the Cordilleras to withstand our
winter climate better than palms of the same species from along the
base-line of the mountains; and so on.
* * * * *
VII. THE ASSUMPTION OF INTERNAL EVOLUTIONARY FORCES
Definite variation was not only postulated in the last decade by Naegeli
and Askenasy, but has also been repeatedly set up in recent years by
various other authors. The Rev. George Henslow, in his book _The Origin of
Species Without the Aid of Natural Selection_, 1894, regards the variations
occurring in the state {83} of nature as always definite and not with
Darwin as indefinite, and meets the objection that modification but not
adaptation to outward conditions of life can be inferred from this fact, by
the bold assumption that it is precisely the outward conditions of life or
the environment which "induces the best fitted to arise." He further
concludes that natural selection has nothing to do with the origin of
species. At the basis of his conviction lies the naturally correct view
that the summation of _accidental_ variations is insufficient for
transforming the species, but that definitely directed variation is
necessary to this end. But concerning the way in which external conditions
are always able to produce the fit variations, he can give us no
information--if I am not mistaken, for the simple reason that such is not
the fact, that the outward conditions only apparently determine the
direction of variations whilst in truth it is the adaptive requirement
itself that produces the useful direction of variation by means of
selectional processes within the germ.
C. Lloyd Morgan also has recently expressed himself in favor of the
necessity of definite variation, though likewise without assigning a basis
for its action, and without being able to show how its efficacy is
compatible with the plain fact of adaptation to the conditions of life. He
seeks to find the origin of variation in "mechanical stresses and chemical
or physical influences," but this conception is too general to be of much
help. He has, in fact, not been able to abandon completely the heredity of
acquired characters.
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