It has been the great service of Roux and his fellow-workers of the
school of “Entwickelungsmechanik,” and of many other students to
whose work we shall refer, to try, as His tried[91] to import into
embryology, wherever possible, the simpler concepts of physics, to
introduce along with them the method of experiment, and to refuse to be
bound by the narrow limitations which such teaching as that of Hertwig
would of necessity impose on the work and the thought and on the whole
philosophy of the biologist.
――――――――――
Before we pass from this general discussion to study some of the
particular phenomena of growth, let me give a single illustration, from
Darwin, of a point of view which is in marked contrast to Haller’s
simple but essentially mathematical conception of Form.
There is a curious passage in the _Origin of Species_[92], where Darwin
is discussing the leading facts of embryology, and in particular Von
Baer’s “law of embryonic resemblance.” Here Darwin says “We are so
much accustomed to see a difference in {58} structure between the
embryo and the adult, that we are tempted to look at this difference
as in some necessary manner contingent on growth. _But there is no
reason why, for instance, the wing of a bat, or the fin of a porpoise,
should not have been sketched out with all their parts in proper
proportion, as soon as any part became visible._” After pointing out
with his habitual care various exceptions, Darwin proceeds to lay down
two general principles, viz. “that slight variations generally appear
at a not very early period of life,” and secondly, that “at whatever
age a variation first appears in the parent, it tends to reappear
at a corresponding age in the offspring.” He then argues that it is
with nature as with the fancier, who does not care what his pigeons
look like in the embryo, so long as the full-grown bird possesses
the desired qualities; and that the process of selection takes place
when the birds or other animals are nearly grown up,—at least on the
part of the breeder, and presumably in nature as a general rule. The
illustration of these principles is set forth as follows; “Let us take
a group of birds, descended from some ancient form and modified through
natural selection for different habits. Then, from the many successive
variations having supervened in the several species at a not very early
age, and having been inherited at a corresponding age, the young will
still resemble each other much more closely than do the adults,—just
as we have seen with the breeds of the pigeon .... Whatever influence
long-continued use or disuse may have had in modifying the limbs or
other parts of any species, this will chiefly or solely have affected
it when nearly mature, when it was compelled to use its full powers
to gain its own living; and the effects thus produced will have been
transmitted to the offspring at a corresponding nearly mature age.
Thus the young will not be modified, or will be modified only in a
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