On the Genesis of SpeciesMivart, St. George Jackson
Religion
On the Genesis of Species
Mivart, St. George Jackson
Evolution (Biology)
{77}
In the cuttle-fishes we find an eye even more completely constructed on the
vertebrate type than is the ear. Sclerotic, retina, choroid, vitreous
humour, lens, aqueous humour, all are present. The correspondence is
wonderfully complete, and there can hardly be any hesitation in saying that
for such an exact, prolonged, and correlated series of similar structures
to have been brought about in two independent instances by merely
indefinite and minute accidental variations, is an improbability which
amounts practically to impossibility. Moreover, we have here again the same
imperfection of the four-gilled cephalopod, as compared with the
two-gilled, and therefore (if the latter proceeded from the former) a
similar indication of a certain comparative rapidity of development.
Finally, and this is perhaps one of the most curious circumstances, the
process of formation appears to have been, at least in some respects, the
same in the eyes of these molluscous animals as in the eyes of vertebrates.
For in these latter the cornea is at first perforated, while different
degrees of perforation of the same part are presented by different adult
cuttle-fishes--large in the calamaries, smaller in the octopods, and
reduced to a minute foramen in the true cuttle-fish sepia.
Some may be disposed to object that the conditions requisite for effecting
vision are so rigid that similar results in all cases must be independently
arrived at. But to this objection it may well be replied that Nature
herself has demonstrated that there is no such necessity as to the details
of the process. For in the higher Annulosa, such as the dragon-fly, we meet
with an eye of an unquestionably very high degree of efficiency, but formed
on a type of structure only remotely comparable with that of the fish or
the cephalopod. The last-named animal might have had an eye as efficient as
that of a vertebrate, but formed on a distinct type, instead of being
another edition, as it were, of the very same structure.
In the beginning of this chapter examples have been given of the very {78}
diverse mode in which similar results have in many instances been arrived
at; on the other hand, we have in the fish and the cephalopod not only the
eye, but at one and the same time the ear also similarly evolved, yet with
complete independence.
Thus it is here contended that the similar and complex structures of both
the highest organs of sense, as developed in the vertebrates on the one
hand, and in the mollusks on the other, present us with residuary phenomena
for which "Natural Selection" alone is quite incompetent to account. And
that these same phenomena must therefore be considered as conclusive
evidence for the action of some other natural law or laws conditioning the
simultaneous and independent evolution of these harmonious and concordant
adaptations.
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