On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
21. —— Scottianus, _Feld._ (Ash Islands).
22. Eurycus Cressida, _Fab._
6. Pacific Islands.
16. Australia.
_Catalogue of Malayan_ PAPILIONIDÆ.
ORNITHOPTERA (Boisd.).
[Illustration:
Fig. 1.
Anal valves of _O. Amphrisius_.
]
The characters in the _larva_ and _pupa_ which have been supposed to
distinguish this genus from PAPILIO are erroneous, or at least do not
exist in all the species. My own observations on _O. Poseidon_ show that
the _larva_ has no “external sheath” to the thoracic tentacles, and that
the suspending thread passes round the pupa, and is not “fastened on
each side to a silky tubercle.” There remain therefore only the
characters of the perfect insect, the most important of which are the
anal valves in the male. These are very large, ovate or rounded,
coriaceous, and not hairy, and are furnished with projecting points or
spines (sometimes very conspicuous) which serve to attach the male more
firmly to the female _in copulâ_. In several species I have observed,
these points or hooks were buried in the protruded anal gland of the
female, and thus effectually prevented the great weight of the insects
causing them to separate upon suddenly taking flight. The great strength
and size of these insects, the thick texture of their wings, their long
curved and stout antennæ, their peculiar form, colour, and distribution,
are the only other characters that separate them from _Papilio_. Though
these may not perhaps be technically sufficient, I think it advisable
and convenient to retain a genus so well known and long established.
_Ornithoptera_ is pre-eminently a Malayan genus, seventeen species
inhabiting the archipelago, one (_Rhadamanthus_, Bd.) India and China,
one (_Darsius_, G. R. Gray) peculiar to Ceylon, one (_Richmondia_, G. R.
Gray) North Australia. _O. Victoriæ_, G. R. Gray, from some island east
of New Guinea, should probably be included in the Malayan list; and
_Æacus_, Felder, from an unknown locality. The following are the
well-established Malayan species.
a. _Priamus_ group.
1. ORNITHOPTERA PRIAMUS, Linnæus.
♂. _Papilio Priamus_, L.; Cram. Pap. Ex. t. 23. f. A, B; Godart,
Enc. Méth. ix. p. 25. _O. Priamus_, Bd. Sp. Gén. Lép. p. 173.
♀. _P. Panthous_, L.; Cram. Pap. Ex. t. 123. f. A, t. 124. f. A.
This may be at once distinguished from all the allied species with which
it has been often confounded—in the _male_, by the more rounded and
deeply scalloped hind wings, with larger black spots and a broader
border, the upper wings with no green on the median nervure or its
branches, and the sooty patch extending only to the second median
nervule; in the _female_, by the very constant and peculiar light
olive-brown colour, the absence of any spots in the discoidal cell of
the upper wings, and the broad shallow scallops of the hinder margin.
_Hab._ Amboyna and Ceram, probably also Bouru (_Wall._).
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