On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
It is a law to which a large proportion of the organic creation would
appear to be subject, that the exuberance of life (not so much,
however, as regards the number of individuals which the various
species may present, as in the grandeur of their size) has reference
to the magnitude of the spot over which it is permitted to range. The
unnatural breeding-in of a single race, which must of necessity happen
unless the intercourse with other varieties of its kind be possible,
has always been attended with effects more or less pernicious; and in
the Annulose tribes I believe that the reduction of space which
geological convulsions have at various epochs brought about, has been
commonly succeeded (_inter alia_) by a reduction of stature in those
species which have been cut off from their fellows. I do not assert
that there are no exceptions to this rule; for counter-influences may
at times prevail (as we shall shortly see), to neutralize the above
tendency. I hold it, however, as an absolute truism, in physics, that
a law without an exception is an anomaly. If, therefore, we were once
to admit the latter to negative the former, no such thing as a law
could exist. Hence it follows, as a corollary (unless, indeed, we are
prepared to endorse that conclusion), _that_ _where there is a law
there must be an exception to it_; and that, consequently, exceptional
cases, if not exceedingly numerous, should never pervert our belief
from an otherwise presumptive truth.
This dwindling-down of size has seldom failed to attract my attention,
more or less, in almost every island which I have hitherto had an
opportunity of exploring: space, however, will not permit me to dwell
upon many instances. I have already adverted to the diminished stature
of _Anthonomus ater_, Mshm, and _Ceutorhynchus contractus_, Mshm, in
Lundy Island,--the first of which scarcely ever reaches, on that rock,
more than half its natural bulk. The late Mr. Holme, of Corpus Christi
College, Oxford, in like manner, captured the common _Calathus
melanocephalus_, Linn., and _Olisthopus rotundatus_, Payk., in
Scilly,--the former of which seldom exceeded two lines, and the latter
two and a half, in length: and he also recorded, that the _Bolitochara
assimilis_, Kby, is invariably smaller in those islands than it is in
the neighbourhood of Penzance[31]. The _Vanessa Callirhoe_, Fabr. (a
geographical analogue of the Red Admiral Butterfly[32], so common in
our own country), is permanently smaller in Porto Santo than it is on
the larger, more luxuriant and varied, and therefore more protected,
island of Madeira proper. And, as regards the _Ptini_ of that group,
so completely are some of them "affected by isolation, and by exposure
to a perpetually stormy atmosphere, that they do not attain half the
bulk on many of the adjacent rocks that they do in the more sheltered
districts of the central mass; and so marvellously is this verified in
Public-domain text, read in full here on John Shaqi.
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