Parasites: A Treatise on the Entozoa of Man and Animals: Including Some Account of the EctozoaCobbold, T. Spencer (Thomas Spencer)
Science
Parasites: A Treatise on the Entozoa of Man and Animals: Including Some Account of the Ectozoa
Cobbold, T. Spencer (Thomas Spencer)
Helminths; Parasites
pronounced developments it is easy to perceive how admirably these
outgrowths are adapted to the necessities of the different species of
parasite to which they are severally referable; and, even in the case of
_Bilharzia_, the trifling amount of anchorage furnished by a projecting
point is not absolutely thrown away. The resistance will also be greater
where the spine is situated a little on one side of the pole of the egg,
which seems to need steadying during the violent struggles of the embryo
to escape from its temporary abode.
[Illustration: FIG. 10.--Free ciliated embryo of _Bilharzia_, with
pear-shaped rudimentary organs below the head. Original.]
[Illustration: FIG. 11.--Two ciliated embryos of _Bilharzia_; showing
sarcode spherules in their interior. That to the left has recently
escaped the shell. Original.]
When any number of ova are removed from the urine and examined, it
will be found that a large proportion of them contain embryos in an
advanced stage of larval growth. The structural appearances presented
by the embryos whilst still in the eggs are remarkably uniform; since,
in all, the yolk appears to have resolved itself into a mass of rounded
sarcode-globules, one or two of these particles being conspicuously
larger than the rest (fig. 12). At this stage, except towards the
cephalic division of the larva, no tendency to differentiation is
perceptible; but some time after the embryo has escaped, one may notice
elongated masses of sarcode formed by the coalescence of the globules.
Whilst still in the egg, one end of the primitive embryonal mass becomes
gradually narrowed, cilia at the same time appearing. This part becomes
the future head, eventually acquiring the form of a cowl. Whatever form
the body of the embryo may display after extrusion from the shell,
the head retains its conical shape, the cone itself being narrowed or
widened only when the larva is subjected to abnormal conditions (fig.
14). Whilst the head is undergoing development within the shell, one,
two, or sometimes three, pyriform masses make their appearance within
the cone; and after the embryo has escaped, these structures become more
marked (fig. 10). The sarcode-globules refract light strongly; and, when
the larva is not compressed in any way, they move freely within the
somatic cavity. In well-developed embryos, whilst still in the egg, the
cilia are observed to clothe every part of the larva except the oral
papilla. This minute nipple-like projection measures about the 1/3000 of
an inch transversely, forming a very simple kind of unarmed proboscis.
When the head of the free embryo is viewed from above, the proboscis
looks like a central ring surrounded by a series of regular folds, which
radiate outwards like the spokes of a wheel. The ridges thus formed
support numerous cilia, these latter projecting at the circumferential
margin of the cephalic cone in such a way as to present the figure of
a star.
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