[Illustration: FIG. 30.--_Tubularia mesembryanthemum._ _A._ Short piece
with reduced hydranth-region. _B._ Piece from distal end of stalk
producing a hydranth without a stalk (see Fig. 27, _D_). _C._ Piece
producing hydranth as outgrowth of end. _C¹._ Later stage of last. _D._
Short piece producing double proboscis (see Fig. 28, _E_).]
in all these cases the basal hydranth develops about twenty-four hours
after the apical one. In the short pieces, however, the two ends develop
at the same time, although the development of all the short pieces,
whatever structures they may produce, whether single or double, is
delayed, and the hydranths may not appear until after the long pieces
have produced their basal hydranths. In these double structures both
ends develop at the same time (Fig. 30, _D_). If we suppose the
influences that start the development of the piece begin first at the
distal end, the region affected will lie so near to the proximal end of
the piece that the development at this end may be hastened, and under
these circumstances the region of new formation will be shared by the
two hydranths. The factors that determine that a larger, partial
structure is formed in preference to a smaller whole one will no doubt
be found to be the same in these double structures and in the single
ones.
_THE INFLUENCE OF THE OLD PARTS ON THE NEW_
One of the most striking and general facts connected with the phenomenon
of regeneration is that the new part that is built up on the exposed
surface is like the part removed. This suggests that an influence of
some sort starts from the old part and changes the part immediately in
contact with it into a structure that completes the old part in that
region. We can imagine that the new part that has been changed in this
way may act on the new part just beyond it, and so step by step the new
part may be differentiated. It is not difficult to show that the
phenomenon is really more complicated than this, and that other factors
are also acting on the new part; but, nevertheless, that the old part
has some such influence is probable. Under certain conditions, however,
this influence may be counteracted by other factors, and something
different from the part removed may be formed. One example of this sort
has already been discussed, namely, that in which after the removal of
much of the anterior end of the earthworm or of a planarian, only the
distal end comes back. Another case is that in which something different
from the part removed is regenerated. If the tip of the eye of the
hermit-crab or of other crustaceans is cut off a new eye is regenerated,
but if the eye-stalk is cut off near its base an antenna-like organ
develops. Herbst has suggested that the presence of the ganglion at the
end of the stalk accounts for the regeneration of a new eye, when only
the tip of the stalk is cut off. In the absence of the ganglion at the
cut-edge the stalk does not produce an eye, but an antenna, as is shown
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