Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
65. _Homogeneous and Allogeneous Nuclei._--The nucleus of the Radiolaria
not only exhibits a similar structure and composition, and suffers similar
modifications to those which are found to occur in the case of other
cell-nuclei, but also to some extent shows very peculiar developmental
forms, which are seldom or never found in other cells. In the first place
the nuclei may be divided into homogeneous and allogeneous, the former are
structureless and consist of a uniform mass of nuclein, whilst the latter
are composed of different substances and show various structural relations.
_Homogeneous_ nuclei, whose whole mass is uniform and exhibits no
structural differentiation, are probably always to be found in the
swarm-spores; in the fully developed Radiolarian body they are found only
in the first legion, SPUMELLARIA, and that both in many Monozoa (especially
small #Sphæroidea# and #Prunoidea#) and in the Polyzoa (or Polycyttaria).
The whole mass of these homogeneous nuclei, which are usually spherical or
ellipsoidal, consists of uniform, perfectly clear and transparent nuclein,
and becomes evenly stained by carmine, hæmatoxyline, &c. They may be
readily distinguished by these means from the clear vacuoles or "hyaline
vesicles," which are evenly distributed in the endoplasm of many
Radiolaria, and may be confused with the former. _Allogeneous_ nuclei,
which are always composed of different parts and often show complicated
structural relations, are found developed in the great majority of
Radiolaria. The most important differentiation exhibited by these secondary
forms is the separation of the nuclear mass into a firm nuclear substance
(caryoplasm) and a fluid nuclear juice (caryolymph). In addition in each
nucleus a nucleolus is visible, and often several or many may be seen (see
§§ 67 to 70).
66. _The Form of the Nucleus._--The nucleus of the Radiolaria shows greater
variations in form and structure than are to be found in the majority of
cell-nuclei; {xxxv}exception must, however, be made in the case of many
animal ovicells, which, in their peculiar form and composition, often
recall large Radiolarian nuclei. With respect to the external shape two
main forms may be distinguished, as primary and secondary. The _primary
form_ of the Radiolarian nucleus is the sphere; it occurs not only in most
swarm-spores, but also in most adult forms belonging to the legion
SPUMELLARIA, and in individual instances in other groups; indeed the nuclei
of most SPUMELLARIA, as also the concentric central capsules in which they
lie, are true geometrical spheres. The _secondary forms_ of the nucleus are
found in the majority of adult Radiolaria, and arise from the primary
spherical forms in various ways, either by the elongation or contraction of
one axis, or by the formation of apophyses or processes. The most important
of these secondary forms are as follows:--
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