Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
B. The fact that in a number of ACANTHARIA the nucleus does not divide
early as in the majority of the legion, but only at a later period, was
first observed by R. Hertwig in a species of #Acanthometra# (_Xiphacantha
serrata_), and a species of #Acanthophracta# (_Phatnaspis
{xxxviii}mülleri_ = _Haliommatidium mülleri_) (_loc. cit._, pp. 11 and
27). This serotinous division of the nucleus seems, however, to be rather
widely spread in both sublegions of the ACANTHARIA; I have found, not
only in the forms above mentioned, but also in several others belonging
to different genera, a single large excentric nucleus, even in those
individuals in which the skeleton was fully developed.
C. The peculiar mode of nuclear budding, by which these small nuclei
arise, appears to proceed in the following manner (Pl. 129). The
vesicular primary nucleus, which, in consequence of the centrogeneous
development of the skeleton protrudes as it grows into irregular lobes
(Pl. 129, fig. 9), assumes a peculiar concavo-convex form, sometimes that
of a hood or dish, sometimes that of a kidney or sausage. The convex
surface is apposed to the capsule-membrane, while the concave is turned
towards the central star of the skeleton (fig. 6). There is now formed at
the centre of the convex surface of the strong, doubly-contoured, nuclear
membrane, a flask-shaped invagination with a narrow neck and expanded
base; the membrane now becomes disposed in peculiar folds, which at the
narrow aperture of invagination appear as folds, but on the expanded body
of the flask take the form of concentric rings, laid closely side by side
(Pl. 129, fig. 10). The convex bottom of the flask, which is directed
towards the concave proximal side of the nucleus, becomes again
invaginated by a central conical apophysis of the enlarged nucleolus,
which is situated between them. Usually the nucleolus has already become
flattened into a lentiform shape, and upon its distal face a conical
apophysis has been developed, which is divisible into a darker proximal
and clearer distal portion. The tip of the latter appears to be in direct
connection with the nuclear membrane at the centre of the base of the
flask-shaped invagination (figs. 6, 10). At this stage of development the
nucleus of the ACANTHARIA generally presents the characteristic form of a
hood-shaped, concavo-convex vesicle, whose radial axis is also the axis
of the flask-shaped distal invagination, and of the depressed conical
nucleolus, which lies between the latter and the concave side of the
nucleus. After this peculiar invagination has persisted for some time in
connection with the enlarged nucleolus, both disappear, and then a
remarkable growth of lobular processes takes place on the concave
proximal side of the hood or kidney-shaped nucleus; from four to eight
knobs of unequal size usually appear, and their thickened wall encloses a
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account