Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The basal apophyses of the sagittal ring are not only of great
morphological importance, because they produce by their union three typical
pairs of basal gates or "collar pores," but also because their
prolongations often appear as typical basal feet. The distal prolongations
of the coracal rods appear in _Semantiscus_ (Pl. 92, figs. 16-18) as two
pectoral feet, those of the scapular rods as two tergal feet, whilst the
opposite prolongations of the basal rod of the sagittal ring appear as two
"sagittal feet" (in front an anterior or sternal, and behind a posterior or
caudal foot). In the typical _Cortiniscus_ (Pl. 92, figs. 11-13) only three
feet are developed; an odd caudal and two paired pectoral feet (compare
above, p. 891). The typical basal ring of the Semantida, with its paired
basal gates (_Semantis_), reappears in the majority of the Coronida,
differing from the former in the development of a second vertical ring,
which lies in the frontal plane (perpendicular to the sagittal ring), and
which we therefore call the frontal ring. In only one small group of the
Coronida the basal ring is absent, namely, in the Zygostephanida, and here
the frontal ring appears in the simplest form, as a complete elliptical
meridian ring, crossing the sagittal ring perpendicularly on the two poles
of the main axis (_Zygostephanus_, Pl. 93, figs. 1-4), Four large lateral
gates between the two rings remain open. This form may be derived directly
from the Stephanida in the following way; from both poles of a simple
sagittal ring there arise two opposite lateral apophyses, which in the
frontal plane become curved one towards the other, and united in the poles
of the transverse axis. The basal apophyses would be the coracal rods. But
it is also possible that _Zygostephanus_ was derived from _Semantis_ by the
loss of the furcular rods.
The three typical rings (or the "dimensive rings") of the #Stephoidea#
appear in their most complete form in the subfamily Trissocyclida (Pl. 93,
figs. 7, 13). Here all three rings are undivided and completely developed
in the three dimensive planes, perpendicular one to another. Between them
there remain eight large open gates; the four superior are the four
"lateral gates" of _Zygostephanus_, the four inferior are the four basal
gates of _Semantrum_. The four latter are originally much smaller than the
four former; but in _Trissocircus_ and _Trissocyclus_ (Pl. 93, figs. 10-12)
they reach the same size. Therefore all eight gates are here of equal form
and similar size, and the basal ring, now a true equatorial ring, divides
the two meridional rings into two equal halves.
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