Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
The clypeiform hypostoma is divided in two fields, one anterior larger,
in the centre elevated in a short, blunt knob, and a posterior, bearing
the two elevated rounded maculæ, just below the semicircular groove
which separates it from the anterior field. These fields are surrounded
by a large border which is posteriorly emarginated through a short
rounded sinus. The surface is entirely covered by small semiglobular
wartlets perforated by a straight pore or duct, which being filled
with a black mineral, probably iron pyrites, gives a peculiar and
characteristic aspect to the sections. In the same manner the whole
mass of the hypostoma is pierced by straight, black lines, also filled
up pores, starting from the outside and ending before attaining the
inner surface.
The maculæ are prominent, separated from the hypostoma through narrow,
distinct grooves, ovate, with the pointed end directed obliquely
outwards, against the margins of the hypostoma. They are covered with
wartlets and leaving only a little oblong spot free, quite smooth, this
being consequently the macula proper. But in a longitudinal section
there is not the least distinction between this macula and the other
hypostoma, only, that it is free of pores.
=Calymmene tuberculata= BRÜNNICH.
Pl. III figs. 8, 9.
We have succeeded to prepare a horizontal section of the macula. This
is oval, perfectly homogenous and without any structure proper, showing
only indistinctly a mottled medley of pale, brownish spots in the clear
mass. There is in both maculæ a small dark, angular spot placed in
their inferior part, fig. 9.
=Centropleura= ANGELIN.
This author has delineated most distinct maculæ in Centr. Steenstrupi
pl. III fig. 4, b. two, small elongate darkish spots, near the
posterior margin. I have not seen any specimen of this trilobite and
have consequently not been able to follow up ANGELIN'S observation.
=Chasmops= M'COY.
Pl. III fig. 10.
In his work on the Russian Phacopidæ FRIEDR. SCHMIDT has given the
figures of Chasmops Eichwaldi and Ch. Wesenbergensis showing, as it
were, patches of black maculæ. But as there is no mention of them, and
as they are quite unlike the macula? which are described here, I am in
doubt of their real nature. We have figured a macula from the hypostoma
of a Chasmops, probably Ch. macroura, from a drift block near Rostock.
It is elongate and narrow, more than four times longer than broad. Its
smooth surface is a little scooped out and no structure is observable.
=Chirurus= BEYRICH.
We find the hypostomic maculæ annotated and delineated as occurring in
the following species.
_Ch. bimucronatus_ SALTER pl. 5, fig. 5 as two narrow elongated pits,
surface seems intact. No mention of them in the letter-press.
_Ch. exsul_ SCHMIDT pl. VI fig. 9. Indications of the oblong, narrow
maculæ.
_Ch. macrophthalmus_ SCHM. pl. VII, figs. 1 c, 2. Faint indications of
narrow pits.
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