Speciation in the Brazilian Spiny RatsMoojen, João
Science
Speciation in the Brazilian Spiny Rats
Moojen, João
Proechimys
_Skull._--Medium in size, not elongated; nasals pointed
posteriorly; supraorbital ridges strong and thick, extending
caudad across anterior third of parietals; zygomatic arches
strong; postorbital process of zygoma involving only squamosal;
incisive foramen 5 by 2.7 mm, oval and extending caudad to plane
approximately 2 mm anterior to premolars; posterior margins of
incisive foramen not forming a ridge; vomerine sheath complete and
with maxillary part slender and threadlike, or incomplete, in
which event, maxillary part not extended enough to join
premaxillary process; mesopterygoid fossa extending forward
as far as centers of third molars; bulla large and more or less
triangular in its peripheral outline.
_Teeth._--Upper molariform teeth with three counterfolds each.
Lower premolar with three counterfolds; lower molars with two
counterfolds.
_Comparisons._--From _P. g. oris_, _arescens_ differs in: Color of
upper parts lighter and more uniform; incisive foramen oval
instead of conspicuously constricted posteriorly; posterior margin
of incisive foramen farther from premolars.
_Remarks._--One specimen from Turí-assú (MN) has been identified by O.
Thomas as "_P. oris_" (his own handwriting is on the label) and the
subspecies is really closely related to _oris_.
_Specimens examined._--Total number, 3, from Brazil, Maranhão, as
follows: _Vitoria do Alto Parnaiba_, Fazenda Inhuma (below Santa
Filomena), 2 (CNHM); Alto da Alegria, Turí-assú, 1 (MN).
=Proechimys guyannensis riparum= subspecies nova
_Type locality._--Manaus, _Manaus_, Amazonas, Brazil. _Type_:
American Museum of Natural History, no. 143018, adult female;
collected 6 March, 1943.
_Range._--Known only from type locality but probably extending
northward and eastward.
_Diagnosis._--Aristiforms wide and stiff; upper parts
Ochraceous-Tawny; incisive foramen short, wide, and oval; vomerine
sheath incomplete; upper molariform teeth and lower premolar with
three counterfolds; lower molars with only two counterfolds.
_Pelage._--_Aristiforms on middorsal region_: Gray basally
gradually blackening toward tip; total length, 18 to 20 mm;
maximum width, 0.9 mm. _Setiforms on middorsal region_: Whitish
basally, gradually blackening toward tip but interrupted by 2 mm
long, Ochraceous-Tawny, subapical zone; total length 20 to 22 mm;
maximum width, 0.04 mm. Some are whitish basally and gradually
become black toward tip with no subapical zone. _Setiforms on
outer thighs_: Whitish basally, gradually becoming gray and then
blackish toward tip but interrupted by Ochraceous-Buff, subapical
zone 3 to 4 mm long; tip not conspicuously dark; total length, 17
to 19 mm; maximum width, 0.03 mm.
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