If we suppose elementary species and varieties originated by sudden
leaps or mutations, then the elementary species have mutated in the line
of progression, some varieties have mutated in the line of
retrogression, while others have diverged from their parental types in a
line of depression, or in the way of repetition. This conception agrees
quite well with the current idea that in the building up of the
vegetable kingdom according to the theory of descent, it is species that
form the links of the chain from the lower forms to the more highly
organized later derivatives. Otherwise expressed, the system is built up
of species, and varieties are only local and lateral, but never of real
importance for the whole structure.
[142] Heretofore we have generally assumed, that varieties differ from
the parent-species in a single character only, or at least that only one
need be considered. We now come to the study of those varieties, which
differ in more than one character. Of these there are two types. In the
first the points of dissimilarity are intimately connected with one
another, in the second they are more or less independent.
The mutually related peculiarities may be termed correlative, and we
therefore speak, in such cases, of correlative variability. This
phenomenon is of the highest importance and is of general occurrence.
But before describing some examples, it is as well to note that in the
lecture on fluctuating variability, cases of a totally different nature
will be dealt with, which unfortunately are designated by the same term.
Such merely fluctuating variations are therefore to be left out of the
present discussion.
The purple thorn-apple, which is considered by some writers as a variety
of the white-flowered species or _Datura Stramonium_, and by others as a
separate species, _D. Tatula_, will serve as an illustration. But as its
distinguishing attributes, as far as we are concerned with them here,
are of the nature described above as characteristic of varietal
peculiarities no objection [143] can be made to our using them as a case
of correlative variability.
The essential character of the purple thornapple lies in the color of
the flowers, which are of a very beautiful pale blue. But this color is
not limited to the corolla. It is also to be seen in the stems and in
the stalks and veins of the leaves, which are stained with a deep
purple, the blue color being added to the original green. Even on the
surface of the leaves it may spread into a purplish hue. On the stems it
is to be met with everywhere, and even the young seedlings show it. This
is of some importance, as the young plants when unfolding their
cotyledons and primary leaves, may be distinguished by this means from
the seedlings of the white flowered species.
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