A familiar feature of the common periwinkle is the lid or stopper
(Operculum), with which the animal can close the mouth of the shell.
This is developed and carried by the outside of the animal's foot. In
the periwinkle and other English molluscs it is comparatively soft and
semi-transparent, and reminds one of a thin slice of horn. In many
tropical molluscs, however, it is hard and shelly. The large tropical
shells named _Turbo_ have massive lids of considerable weight. These
shells, which are nearly allied to the pearly Top-shells (_Trochus_) of
the English shores, are sold as ornaments, the outer coat of the shell
being partly scraped off to show the inner coat of pearl: it is rarely,
however, that the purchaser obtains a lid, or even knows that the
creature had one. The reverse is the case with some of the smaller
kinds, the lids of which, being brightly coloured, are imported without
the shell, and sometimes set as articles of jewellery. Some of these are
of a bright green hue.
While the lids of the Holostomata are rounded in shape, those that
belong to the Siphonostomatous shells are necessarily more or less
modified so as to fit the mouth of the shell, and are consequently oval
or even claw-like in shape. The Sting-winkle already spoken of, the
common small whelk, _Purpura lapillus_, and the large whelk, _Buccinum
undatum_, are common shell-fish in which the elongated lid may be
studied. The lid is not, however, like the tongue-ribbon, an essential
feature of the structure of every univalve mollusc.[D] Not only are
there special instances in which it is greatly modified, but also
there are whole groups of univalve molluscs in which it is absent.
[D] There are one or two exceptional cases of gasteropod molluscs that
have no tongue-ribbon. The majority of these are parasitic forms, which
can get their food without the trouble of filing it down.
A curious suggestion has been made with regard to the lids of univalve
shell-fish; namely, that the snapping to of the lid is capable of
producing a sound, which may perhaps be audible at a distance under the
water. Various molluscs have been credited with producing sounds, either
by muscular movements or by the grating of the shell as the animal
walks. The common Tortoise-shell Snail, _Helix aspersa_, sometimes
makes a most alarming noise when crawling over a window. It has been
disputed whether the sounds thus made are produced by the grating of the
creature's tongue-ribbon on the glass, as it files off small particles
of algae and vegetable moulds, which are invisible to our eyes: or
whether they are sounds due to suction of the muscular surfaces, such as
may be produced by drawing a wet finger across glass. The noise, however
produced, is, as I can testify from experience, sufficiently loud and
weird to be very startling, if heard in the dead of night.
Public-domain text, read in full here on John Shaqi.
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