We have here therefore another case like that which we met with among
the Rhopalocerous Lepidoptera, in which the imagines appear to be
capable of being formed into a higher group than the larvæ, because the
former live under conditions of life which are on the whole similar
whilst the latter live under very divergent conditions.
The old division of the Hymenoptera into two sub-orders has certainly
been abandoned in the later zoological text-books; they are now divided
into three:--saw-flies, parasitic, and aculeate Hymenoptera; but even
this arrangement has been adopted with reference to the different
structure of the larvæ. Whether this system is better than the older,
_i.e._ whether it better expresses the _genealogical_ relationship, I
will not now stop to investigate.[201]
DIPTERA.
The imagines of the Diptera (_genuina_), with the exception of the
_Aphaniptera_ and _Pupipara_, agree in all their chief characters,
such as the number and structure of the wings, the number and joints
of the legs, the peculiar formation of the thorax (fusion of the three
segments);[202] and even the structure of the mouth organs varies only
within narrow limits. This is in accordance with their mode of life,
which is very uniform in its main features: all the true Diptera live
in the light, moving chiefly by means of flight, but having also the
power of running; all those which take food in the imago condition
feed upon fluids. Their larvæ, on the other hand, are formed on two
essentially different types, the one--which I shall designate as the
gnat-type--possessing a horny head with eyes, three pairs of jaws,
and long or short antennæ, together with a 12- or 13-segmented body,
which is never provided with the three typical pairs of thoracic legs,
but frequently has the so-called abdominal legs on the first and last
segments. The other Dipterous larvæ are maggot-shaped and without a
horny head, or in fact without any head, since the first segment, the
homologue of the head, can in no case be distinguished through its
being larger than the others; it is on the contrary much smaller. The
typical insect mouth-parts are entirely absent, being replaced by a
variously formed and quite peculiar arrangement of hooks situated on
the mouth and capable of protrusion. Never more than eleven segments
are present besides the first, which is destitute of eyes; neither are
abdominal legs ever developed.
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