Further facts can be adduced in support of the view that in these
processes it is the constitution of the species, and not the external
agency (warmth), which plays the chief part. The latter, as Darwin
has strikingly expressed it, rather performs the function of the
spark which ignites a combustible substance, whilst the character of
the combustion depends upon the nature of the explosive material.
Were this not the case, increased warmth would always change a given
colour[29] in the same manner in all butterflies, and would therefore
always give rise to the production of the same colour. But this does
not occur; _Polyommatus Phlæas_, for example, becoming black in the
south, whilst the red-brown _Vanessa Urticæ_ becomes black in high
northern latitudes, and many other cases well known to entomologists
might be adduced.[30] It indeed appears that species of similar
physical constitution, i.e., nearly allied species, under similar
climatic influences, change in an analogous manner. A beautiful example
of this is furnished by our _Pierinæ_. Most of the species display
seasonal dimorphism; as, for instance, _Pieris Brassicæ_, _Rapæ_,
_Napi_, _Krueperi_, and _Daplidice_, _Euchloe Belia_ and _Belemia_,
and _Leucophasia Sinapis_, in all of which the difference between the
winter and the summer forms is of a precisely similar nature. The
former are characterized by a strong black dusting of the base of the
wings, and by a blackish or green sprinkling of scales on the underside
of the hind wings, while the latter have intensely black tips to the
wings, and frequently also spots on the fore-wings.
Nothing can prove more strikingly, however, that in such cases
everything depends upon the physical constitution, than the fact that
in the same species the males become changed in a different manner to
the females. The parent-form of _Pieris Napi_ (var. _Bryoniæ_) offers
an example. In all the _Pierinæ_ secondary sexual differences are
found, the males being differently marked to the females; the species
are thus sexually dimorphic. Now the male of the Alpine and Polar var.
_Bryoniæ_, which I conceive to be the ancestral form, is scarcely to
be distinguished, as has already been mentioned, from the male of our
German winter form (_P. Napi_, var. _Vernalis_), whilst the female
differs considerably.[31] The gradual climatic change which transformed
the parent-form _Bryoniæ_ into _Napi_ has therefore exerted a much
greater effect on the female than on the male. The external action on
the two sexes was exactly the same, but the response of the organism
was different, and the cause of the difference can only be sought for
in the fine differences of physical constitution which distinguish the
male from the female. If we are unable to define these differences
precisely, we may nevertheless safely conclude from such observations
that they exist.
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