Now if, returning to questions previously solved, the alternating
action of cold in winter and warmth in summer leads to the production
of a winter and summer form, according to the law of cyclical heredity,
the question still remains: why do we not find seasonal dimorphism in
all polygoneutic butterflies?
We might at first suppose that all species are not equally sensitive
to the influence of temperature: indeed, the various amounts
of difference between the winter and summer forms in different
species would certainly show the existence of different degrees of
sensitiveness to the modifying action of temperature. But even this
does not furnish an explanation, since there are butterflies which
produce two perfectly similar[34] generations wherever they occur,
and which, nevertheless, appear in different climates as climatic
varieties. This is the case with _Pararga Ægeria_ (Fig. 23, Plate
II.), the southern variety of which, _Meione_ (Fig. 24, Plate II.),
is connected with it by an intermediate form from the Ligurian coast.
This species possesses, therefore, a decided power of responding to
the influence of temperature, and yet no distinction has taken place
between the summer and the winter form. We can thus only attribute
this different deportment to a different kind of heredity; and we
may therefore plainly state, that changes produced by alternation
of climate are not always inherited _alternatingly_, i.e. by the
corresponding generations, but sometimes _continuously_, appearing
in every generation, and never remaining latent. The causes which
determine why, in a particular case, the one or the other form of
inheritance prevails, can be only innate, i.e. they lie in the organism
itself, and there is as little to be said upon their precise nature as
upon that of any other process of heredity. In a similar manner Darwin
admits a kind of double inheritance with respect to characters produced
by sexual selection; in one form these characters remain limited to the
sex which first acquired them, in the other form they are inherited by
both sexes, without it being apparent why, in any particular case, the
one or the other form of heredity should take place.
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