first place, in different species there are very different degrees of
difference between the summer and winter forms; and secondly, many
digoneutic species are still monomorphic in Germany, first becoming
seasonally dimorphic in Southern Europe. This is the case with _P.
Machaon_ and _P. Podalirius_, as already mentioned, and likewise with
_Polyommatus Phlæas_. Zeller in 1846-47, during his journey in Italy,
recognized as seasonally dimorphic in a small degree a large number of
diurnal Lepidoptera which are not so in our climate.[35]
In a similar manner the appearance of seasonal dimorphism in species
which, like _Plebeius Amyntas_, do not hibernate as pupæ, but as
caterpillars, can be simply explained by supposing that the winter
generation was the primary form, and that the increase in the summer
temperature since the glacial period was sufficient to cause this
particular species to become changed by the gradual interpolation
of a second generation. The dimorphism of _P. Amyntas_ can,
nevertheless, be explained in another manner. Thus, there may have
been a disturbance of the period of development in the manner already
indicated, the species which formerly hibernated in the pupal stage
becoming subsequently disturbed in its course of development by the
interpolation of a summer generation, and hibernating in consequence
in the caterpillar state. Under these circumstances we must regard the
present winter form (var. _Polysperchon_) as having been established
under the influence of a winter climate, this form, since the supposed
disturbance in its development, having had no reason to become changed,
the spring temperature under which its pupation now takes place not
being sufficiently high. The interpolated second generation on the
other hand, the pupal period of which falls in the height of summer,
may easily have become formed into a summer variety.
This latter explanation agrees precisely with the former, both starting
with the assumption that in the present case, as in that of _A. Levana_
and the _Pierinæ_, the winter form is the primary one, so that the
dimorphism proceeds from the said winter form and does not originate
the winter but the summer form, as will be explained. Whether the
winter form has been produced by the action of the winter or spring
temperature is immaterial in judging single cases, inasmuch as we are
not in a position to state what temperature is necessary to cause any
particular species to become transformed.
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