The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
The exopodite on this same specimen was evidently cut diagonally
across near the setiferous edge, showing a section through the shaft
and the bases of seven setæ (fig. 18). This section is so exactly what
would be obtained by cutting similarly an exopodite of either Neolenus
or _Triarthrus_ that it should in itself dispose of the
"spiral-exopodite" theory.
Several sections have already been illustrated showing sections across
the setæ of the exopodites (pl. 3, figs. 4-6, 1881; pl. 27, figs. 3,
4, 9, 1918), and similar sections are not uncommon. Only a very few,
however, show sections in the plane of the exopodite. If only No. 111,
described above, were known, it would be inferred that the exopodite
had a slender shaft as in _Calymene_, but another good slice, No. 80
(fig. 12, ante) shows that the blade was rather broad, though not so
broad as in Neolenus. The other specimen is No. 22, which has already
been discussed. The thoracic exopodite of this specimen has been very
incorrectly figured by Walcott, as it shows no such palmate shaft as
he has indicated, but a long blade-like one is outlined, though its
entire width is not actually shown.
_Pygidial Appendages._
Sections 14 and 18 (pl. 2, figs. 4, 8, 1881) prove the presence under
the pygidium of three pairs of appendages, the coxopodites and
fragments of endopodites of which are shown. Nothing is known of the
exopodites.
_Relation of Hypostoma to Cephalon._
In Ceraurus the body portion and posterior end of the hypostoma are
roughly oval, about as wide as the glabella at its broadest part, and
the posterior edge extends back to within 0.5 to 1 mm. of the neck
furrow. The posterior pair of appendifers are behind the hypostoma,
while the second pair are in front of its posterior end but escape
being covered by it on account of its oval shape. At the anterior end
the hypostoma is widened by the presence of two side lappets which
extend beyond the boundaries of the glabella. In both Ceraurus and
Cheirurus the anterior edge of the hypostoma fits against the doublure
at the anterior margin of the head and the epistoma is either entirely
absent or is so narrow as not to be seen in specimens in the ordinary
state of preservation. A section across the cephalon of _Ceraurus
pleurexanthemus_ at the horizon of the eyes shows the sides of the
hypostoma fitting closely against the sides of the glabella (Walcott's
pl. 1, fig. 1). Further back on the head it is not in contact with the
dorsal test, and the gnathobases extend beneath it.
Restoration of _Ceraurus pleurexanthemus_. (pl. 11; text fig. 19.)
The restoration of the appendages of _Ceraurus pleurexanthemus_ is a
tentative one, based upon a careful study of the translucent sections
prepared by Doctor Walcott. In no case among these sections is the
actual test of any appendage preserved, and the real form of each part
is generally obscured by the crystallization of the calcite which
fills the spaces formerly occupied by animal matter.
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